materials from different regions of Russia are encoded as: FS RU CU UR WS SS MS CY NE NF KM AM SH KU
Bolboceratinae
Odontaeus armiger Scopoli, 1772 RU
Crimea
Hungary
Eubolbitus radoszkowskii (Solsky, 1876) Kazakhstan
Kyrghyzstan
Bolbocerodema zonatum Nikolajev, 1973
AM
Bolbocerodema nigroplagiatum (Waterhouse, 1875) Japan
Eucanthus lazarus (Fabricius, 1775) USA
Geotrupinae
Geotrupini
Ceratophius polyceros Pallas, 1771 RU
WS
Ceratophius dauricus Jekel, 1865 SS Mongolia
Geotrupes (Geotrupes) mutator Marsham, 1802 RU
CU UR Azerbaijan
Geotrupes (Geotrupes) spiniger Marsham, 1802 Armenia
Azerbaijan Italy
Geotrupes (Geotrupes) niger Marsham, 1802
Italia
Geotrupes (Geotrupes) stercorarius Linnaeus, 1758 RU
Geotrupes (Geotrupes) baicalicus Reitter, 1893 RU
UR WS SS Hungary Kazakhstan
Geotrupes (Geotrupes) jakovlevi Semenov, 1891 Kazakhstan
Kyrghyzstan
Geotrupes (Geotrupes) amoenus G.Jacobson, 1893
SS AM
Geotrupes (Geotrupes) olgae Olsoufjev, 1918
CU Ukraine Turkey Georgia
Geotrupes (Anoplotrupes) stercorosus Scriba, 1791 RU
UR WS SS Italy Finland Belarus Ukraine
Geotrupes (Trypocopris) vernalis Linnaeus, 1758
Slovakia Hungary Yugoslavia
Geotrupes (Trypocopris) fulgidus Motschulsky, 1845
Bulgaria
Geotrupes (Trypocopris) caspicus Motschulsky, 1845 RU
Geotrupes (Trypocopris) inermis Menetries, 1832
Georgia
Geotrupes (Glyptogeotrupes) impressus Gebler, 1841 Kazakhstan
Kyrghyzstan
Geotrupes (Glyptogeotrupes) geminatus Gene, 1839
France(Corse)
Geotrupes (Phelotrupes) laevistriatus Motschulsky, 1857
KU
Geotrupes (Phelotrupes) auratus Motschulsky, 1857
AM KU Japan
Mycotrupes sp. USA
Lethrinae
Lethrus (Ceratodirus) karelini Gebler, 1845 Kazakhstan
Kyrghyzstan
Lethrus (Ceratodirus) longimanus Fischer von Waldheim, 1821 Kazakhstan
Lethrus (Ceratodirus) borealis Nikolajev, 1973 paratype!
Kazakhstan
Lethrus (Ceratodirus) gladiator Reitter, 1897 Kazakhstan
Lethrus (Abrognathus) pygmaeus Ballion, 1870
Tadzhikistan Uzbekistan
Lethrus (Abrognathus) tuberculifrons Ballion, 1870 Kazakhstan
Tadzhikistan
Lethrus (Lethrus) scoparius Fischer, 1831
Lethrus (Lethrus) scoparius scoparius Fischer, 1831 Kazakhstan
Lethrus (Lethrus) scoparius laevigatus Ballion, 1870
Uzbekistan
Lethrus (Lethrus) ?politus Solsky, 1876
Uzbekistan
Lethrus (Lethrus) superbus Kraatz, 1882
Uzbekistan Turkmenistan
Lethrus (Lethrus) inermis Reitter, 1897
Tadzhikistan
Lethrus (Lethrus) geminatus Kraatz, 1882
Uzbekistan
Lethrus (Lethrus) apterus Laxman, 1870 RU
Ukraine
Lethrus (Lethrus) hauseri Reitter, 1894 Kazakhstan
Uzbekistan
Lethrus (Lethrus) sieversi D.Koshantschikov, 1894 Turkmenistan
Lethrus (Lethrus) nikolajevi Medvedev, 1971
Tadzhikistan
Lethrus (Lethrus) arnoldii Nikolajev, 1987
Uzbekistan
Lethrus (Lethrus) glaber Medvedev, 1962
Tadzhikistan
Lethrus (Lethrus) turkestanicus Ballion, 1870 (=aenescens
Fairmaiere, 1892) Kazakhstan Uzbekistan
Lethrus (Scelolethrus) substriatus Kraatz,1883 (=turcomanicus
Semenov, 1894) Turkmenistan
Lethrus (Scelolethrus) spinimanus B.Jakovlev, 1892
Turkmenistan
Lethrus (Scelolethrus) rosmarus Ballion, 1870 Kazakhstan
Uzbekistan Turkmenistan
Lethrus (Scelolethrus) sulcatus Kraatz, 1883
Turkmenistan
Lethrus (Scelolethrus) tekkensis Kral et Olexa, 1996
Turkmenistan
Lethrus (Scelolethrus) costatus Semenov, 1894
Tadzhikistan
Lethrus (Furcilethrus) frantsevichi Nikolajev, 1979
Tadzhikistan
Lethrus (Furcilethrus) microbuccis Ballion, 1870
Tadzhikistan
Lethrus (Heteroplistodus) cicatricosus Reitter, 1890
Turkmenistan
Lethrus (Heteroplistodus) bulbocerus Fischer, 1845 (=jakovlevi
Semenov, 1892)
Lethrus (Heteroplistodus) bulbocerus tarbagataicus Nikolajev, 2001 Kazakhstan
Lethrus (Heteroplistodus) serridens Nikolajev, 1971 Tadzhikistan
Lethrus (Heteroplistodus) tschitscherini Semenov, 1894
Kazakhstan
Ochodaeinae
Codocera ferruginea Eschscholtz, 1818 RU
WS SS AM Finland Hungary Kazakhstan Mongolia
Ochodaeus cornifrons Solsky, 1876 Turkmenistan
Glaphyrinae
Amphicoma (Solskiola) analis Solsky, 1876 Turkmenistan
Uzbekistan Tadzhikistan
Amphicoma (Eulasia) regeli Ballion, 1878 Turkmenistan
Uzbekistan
Amphicoma (Eulasia) kuschakevitschi Ballion, 1871
Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan
Tadzhikistan
Amphicoma (Eulasia) bombyliformis (Pallas, 1781)
Kazakhstan Turkmenistan
Amphicoma (Eulasia) arctos (Pallas, 1781)
CU Georgia Armenia
Amphicoma (Pygopleurus) vulpes (Fabricius, 1781) RU
Amphicoma (Pygopleurus) psilotrichia Faldermann, 1835
Amphicoma (Pygopleurus) banghaasi Reitter, 1895 Turkmenistan
Tadzhikistan
Glaphyrus micans Faldermann, 1835 Armenia
Glaphyrus turkestanicus Semenov, 1889
Tadzhikistan
Glaphyrus oxypterus (Pallas, 1771) Kazakhstan