materials from different regions of Russia are encoded as: FS RU CU UR WS SS MS CY NE NF KM AM SH KU

Hybosorinae
Hybosorus illigeri Reiche, 1853 Turkmenistan Tadzhikistan

Scarabaeinae
Scarabaeus sacer Linnaeus, 1758 RU Azerbaijan Turkmenistan
Scarabaeus semipunctatus Fabricius, 1792 Spain
Scarabaeus pius Illiger, 1803 CU Armenia Turkmenistan Tadzhikistan
Scarabaeus typhon Fischer, 1822 RU Kazakhstan Turkmenistan
Scarabaeus carinatus Gebler, 1841 Uzbekistan Turkmenistan Tadzhikistan
Scarabaeus babori Balthasar, 1934 Uzbekistan Turkmenistan
Scarabaeus acuticollis Motschulsky, 1849 Turkmenistan Tadzhikistan
Scarabaeus armeniacus Menetries, 1832 CU Georgia
Scarabaeus transcaspicus Stolfa, 1938 Turkmenistan
Gymnopleurus flagellatus (Fabricius, 1787) CU WS Azerbaijan Kazakhstan Turkmenistan
Gymnopleurus mopsus Pallas, 1781 RU SS Ukraine Kazakhstan Turkmenistan
Gymnopleurus aciculatus Gebler, 1845 Kazakhstan Uzbekistan Turkmenistan Tadzhikistan
Gymnopleurus sturmi MacLeay, 1821 Bulgaria
Gymnopleurus geoffroyi Fuessly, 1875 RU UR
Gymnopleurus smaragdinus Fahraeus in Boheman, 1857 Uganda
Gymnopleurus sinuatus (Olivier, 1789) China(Taiwan)

Coprinae
Sisyphus schaefferi Linnaeus, 1758
Sisyphus schaefferi boschniaki Fischer von Waldhaim, 1823 RU UR Bulgaria Crimea Armenia Georgia Turkmenistan
Sisyphus schaefferi morio Arrow, 1909 AM
Synapsis tmolus Fischer, 1821 Kazakhstan Kyrghyzstan Turkmenistan Tadzhikistan
Copris lunaris (Linnaeus, 1758) RU CU UR Latvia Crimea Kazakhstan Kyrghyzstan Turkmenistan
Copris hispanus (Linnaeus, 1764) Ukraine Uzbekistan Turkmenistan Tadzhikistan
Copris ochus Motschulsky, 1860 AM
Copris pecuarius Lewis, 1884 AM
Copris integer Reiche, 1847 Rwanda
Copris orion Klug, 1835 Rwanda
Copris magicus Harold,  Vietnam
Copris (Microcopris) reflexus F. Vietnam
Copris (Microcopris) propinquus Fels.  Vietnam
Heliocopris hamadryas (Fabricius, 1775) Uganda
Onthophagus (Euonthophagus) amyntas (Olivier, 1789) Kazakhstan Kyrghyzstan Uzbekistan Turkmenistan Tadzhikistan
Onthophagus (Euonthophagus) gibbosus (Scriba, 1790) Azerbaijan Kazakhstan Kyrghyzstan Uzbekistan Turkmenistan Tadzhikistan
Onthophagus (Euonthophagus) sulcicollis Reitter, 1892 Turkmenistan Tadzhikistan Afghanistan
Onthophagus (Euonthophagus) crocatus Mulsant, 1873 Algeria
Onthophagus (Euonthophagus) dorbignyi Olsoufieff, 1900 Afghanistan
Onthophagus (Altonthophagus) sibiricus Harold, 1877 Kazakhstan Kyrghyzstan Tadzhikistan
Onthophagus (Altonthophagus) marmotae Kabakov, 1990 Tadzhikistan
Onthophagus (Altonthophagus) uniformis Heyden, 1886 AM
Onthophagus (Relictonthophagus) punctator Reitter, 1893 SS AM
Onthophagus (Gibbonthophagus) atripennis Waterhouse, 1875 AM
Onthophagus (Gibbonthophagus) fodiens Waterhouse, 1875 AM
Onthophagus (Gibbonthophagus) limbatus (Herbst) Philippines
Onthophagus (Phanaeomorphus) japonicus Harold, 1874 AM
Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) RU CU UR Georgia Armenia Azerbaijan Turkmenistan
Onthophagus (Colobonthophagus) transcaspicus Koenig, 1888 Turkmenistan
Onthophagus (Colobonthophagus) lunatus Harold, 1868 Vietnam
Onthophagus (Onthophagus) taurus (Schreber, 1759) RU CU Ukraine Crimea Georgia Iran Turkmenistan Uzbekistan
Onthophagus (Onthophagus) illyricus {Scopoli, 1763) Crimea Azerbaijan
Onthophagus (Onthophagus) bivertex Heyden, 1887 AM
Onthophagus (Palaeonthophagus) vitulus (Fabricius, 1776) RU UR WS Kazakhstan Bulgaria
Onthophagus (Palaeonthophagus) citellorum Medvedev, 1965 RU
Onthophagus (Palaeonthophagus) quadrinodus Reitter, 1896 Uzbekistan
Onthophagus (Palaeonthophagus) gibbulus (Pallas, 1781) RU CU UR WS SS MS AM Crimea Kazakhstan Kyrghyzstan
Onthophagus (Palaeonthophagus) laticornis Gebler, 1823 SS
Onthophagus (Palaeonthophagus) nuchicornis (Linnaeus, 1758) RU UR WS SS Kazakhstan
Onthophagus (Palaeonthophagus) vacca (Linnaeus, 1767) RU CU UR Moldova Azerbaijan Turkmenistan
Onthophagus (Palaeonthophagus) lemur (Fabricius, 1781) CU Moldova Turkmenistan
Onthophagus (Palaeonthophagus) fissicornis Krynisky, 1834 CU Armenia Azerbaijan Turkmenistan
Onthophagus (Palaeonthophagus) suturellus Brulle, 1832 RU Azerbaijan
Onthophagus (Palaeonthophagus) conspersus Reitter, 1892 Tadzhikistan Afghanistan
Onthophagus (Palaeonthophagus) marginalis Gebler, 1817 WS SS Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan Mongolia
Onthophagus (Palaeonthophagus) speculifer Solsky, 1876 Kazakhstan Kyrghyzstan Turkmenistan
Onthophagus (Palaeonthophagus) basipusctulatus Heyden, 1889 Turkmenistan
Onthophagus (Palaeonthophagus) pygargus Motschulsky, 1845 Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan Tadzhikistan
Onthophagus (Palaeonthophagus) ovatus (Linnaeus, 1762) RU CU Bulgaria Crimea
Onthophagus (Palaeonthophagus) semicornis (Panzer, 1798) RU UR SS Kazakhstan Turkmenistan
Onthophagus (Palaeonthophagus) medvedevi Kabakov, 1982 Uzbekistan
Onthophagus (Palaeonthophagus) vlasovi Medvedev, 1958 Kazakhstan Turkmenistan
Onthophagus (Palaeonthophagus) glasunovi Koshantschikov, 1894 Tadzhikistan
Onthophagus (Palaeonthophagus) finschi Harold, 1877 Kazakhstan Kyrghyzstan Turkmenistan
Onthophagus (Palaeonthophagus) afghanus Petrovitz, 1961 Tadzhikistan
Onthophagus (Palaeonthophagus) trispinus Reitter, 1892 Turkmenistan Tadzhikistan
Onthophagus (Palaeonthophagus) leucostigma Stevens, 1811 UR Kazakhstan Turkmenistan Uzbekistan Tadzhikistan
Onthophagus (Palaeonthophagus) haroldi Ballion, 1871 Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan Tadzhikistan
Onthophagus (Palaeonthophagus) truchmenus Kolenati, 1846 Azerbaijan Turkmenistan
Onthophagus (Palaeonthophagus) ruficapillus Brulle, 1832 CU Armenia Azerbaijan
Onthophagus (Palaeonthophagus) grossepunctatus J.Muller, 1906 CU
Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781) Azerbaijan
Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790) RU CU UR WS SS Hungary Moldova Ukraine Georgia Armenia Kazakhstan Turkmenistan
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783) Moldova Georgia
Onthophagus (Palaeonthophagus) angorensis Petrovitz Turkmenistan
Onthophagus (Palaeonthophagus) pseudocaccobius Reitter, 1889 Turkmenistan
Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 SS AM
Onthophagus (Palaeonthophagus) scabriusculus Harold, 1873 RU WS SS AM
Onthophagus (Palaeonthophagus) viridis Menetries, 1832 Azerbaijan
Onthophagus (Palaeonthophagus) clitellifer Reitter, 1894 SS
Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893 Bulgaria
Onthophagus (Parascatonomus) multicifrons (Endrodi, 1973) Vietnam
Onthophagus cuniculus Macley Australia
Diastellopalpus lamellicollis Qued. Rwanda
Caccobius (Caccobius) jessoensis Harold, 1867 KU
Caccobius (Caccobius) brevis Waterhouse, 1875 AM
Caccobius (Caccobius) schreberi (Linnaeus, 1767) RU CU UR Ukraine Crimea Georgia Turkmenistan
Caccobius (Caccobius) histeroides (Menetries, 1832) RU CU Crimea
Caccobius (Caccobius) denticollis Harold, 1897 Afghanistan
Caccobius (Caccobius) hymalayanus Jekel, 1772 Afghanistan
Caccobius (Caccophilus) christophi Harold, 1879 AM
Caccobius (Caccophilus) sordidus Harold, 1886 AM
Caccobius (Caccophilus) kelleri (Olsoufieff, 1907) AM
Chironitis pamphilus Menetries, 1849 Uzbekistan Turkmenistan
Chironitis eumenes (Motschulsky, 1859) Ukraine Kazakhstan
Chironitis haroldi Ballion, 1870 Kazakhstan Uzbekistan Turkmenistan
Chironitis moeris Pallas, 1781 RU Kazakhstan
Chironitis sterculius Ballion, 1870 Uzbekistan Turkmenistan
Chironitis klapperichi Ballion, 1870 Iran
Onitis damoetas Steven, 1806 Georgia
Onitis humerosus (Pallas, 1771) Kazakhstan Uzbekistan Turkmenistan
Onitis crenatus Reiche, 1850 Rwanda
Onitis falcatus (Wulfen, 1789) Thailand
Onitis subopacus Arrow, 1931 Vietnam
Euoniticellus fulvus (Goeze, 1777) CU UR WS Moldova Ukraine Crimea Georgia Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan Tadzhikistan
Euoniticellus pallipes (Fabricius, 1781) RU CU Israel Turkmenistan Tadzhikistan

Aphodiinae
Aegialiini
Aegialia (Aegialia) comis (Levis, 1895) AM
Aegialia (Psammoporus) abdita (Nikritin, 1975) SS
Aegialia (Psammoporus) friebi Balthasar, 1935 SH
Aegialia (Psammoporus) kamtschatica Motschulsky, 1860 AM SH KM
Aegialia (Psammoporus) sabuleti Panzer, 1797 RU
Aegialia (Psammoporus) cylindrica (Eschscholtz, 1822) USA
Eremazus unistriatus Mulsant, 1851 Turkmenistan
Eremazus cribratus Semenov, 1893 Turkmenistan

Psammodini
Diastictus vulneratus Sturm, 1805 WS SS
Psammodius asper (Fabricius, 1775) SS
Psammodius (Rhyssemus) orientalis (Mulsant et Codart, 1875) Kazakhstan Turkmenistan
Psammodius (Rhyssemus) germanus (Linnaeus, 1767) CU WS SS Georgia Turkmenistan Tadzhikistan
Psammodius (Rhyssemus) laevipennis Costa, 1744 Ukraine
Psammodius (Pararhyssemus) coluber Mayet, 1787 Turkmenistan Afghanistan
Pleurophorus variolosus (Kolenati, 1846) RU Ukraine Kazakhstan Turkmenistan Tadzhikistan
Pleurophorus caesus Creutzer, 1796 CU Moldova Georgia Turkmenistan Tadzhikistan
Pleurophorus apicipennis Reitter, 1892  Kazakhstan Turkmenistan Tadzhikistan

Thinorycterini
Thinorycter chlamydatus Semenov et Reichardt, 1925 Turkmenistan

Aphodiini
Turanella latevittis (Reitter, 1887) Turkmenistan
Oxyomus silvestris (Scopoli, 1763) RU CU Georgia
Heptaulacus testudinarius (Fabricius, 1775) CU
Heptaulacus sus (Herbst, 1783) RU UR WS Armenia Hugary
Heptaulacus carinatus (Germar, 1824) UR WS SS Kazakhstan
Heptaulacus villosus (Gyllenhal, 1806) SS
Aphodius (Colobopterus) erraticus (Linnaeus, 1758) RU UR WS SS MS Ukraine Kazakhstan Turkmenistan Uzbekistan
Aphodius (Colobopterus) propraetor Balthasar, 1932 AM
Aphodius (Colobopterus) quadratus Reiche, 1847 KU
Aphodius (Coprimorphus) scrutator (Herbst, 1789) Italy Hugary Crimea
Aphodius (Eupleurus) subterranius (Linnaeus, 1758) RU CU UR WS SS CY Kazakhstan Turkmenistan
Aphodius (Eupleurus) antiquus Faldermann, 1835 SS Mongolia
Aphodius (Teuchestes) brachysomus Solsky, 1874 KU
Aphodius (Teuchestes) fossor (Linnaeus, 1758) RU CU UR WS SS Ukraine Crimea Kazakhstan
Aphodius (Otophorus) haemorrhoidalis (Linnaeus, 1758) RU UR WS SS AM SH KU Italy Ukraine Kazakhstan
Aphodius (Lunaphodius) lunifer Solsky, 1876 Turkmenistan Tadzhikistan
Aphodius (Ammoecius) brevis Erichson, 1848 RU UR WS Kazakhstan
Aphodius (Aparammoecis) isaburoi Nakane, 1956 Japan
Aphodius (Bodilus) hydrohoeris (Fabricius, 1798) France
Aphodius (Bodilus) sordidus (Fabricius, 1775) WS SS Mongolia
Aphodius (Bodilus) rufus (Moll, 1782) UR Kazakhstan Hugary
Aphodius (Bodilus) ictericus (Laicharting, 1781) RU CU WS SS Hungary Crimea Kazakhstan
Aphodius (Bodilus) punctipennis Erichson, 1848 WS Kazakhstan Turkmenistan
Aphodius (Bodilus) longeciliatus Reitter, 1887 SS
Aphodius (Bodilus) lugens Creutzer, 1799 RU CU Ukraine Crimea Kazakhstan Kyrghyzstan Turkmenistan Tadzhikistan
Aphodius (Bodilus) sordescens Harold, 1869 WS SS Kazakhstan Mongolia
Aphodius (Bodilus) gregarius Harold, 1871 RU Kazakhstan Mongolia
Aphodius (Bodilus) digitalis D.Koshantschikov, 1894 Turkmenistan
Aphodius (Bodilus) hastatus Reitter, 1892 Kazakhstan
Aphodius (Bodilus) strigimargo Reitter, 1892 Turkmenistan
Aphodius (Bodilus) nigriventris Reitter, 1892 Kazakhstan
Aphodius (Acanthbodilus) immundus Creutzer, 1799 CU UR WS SS Kazakhstan Turkmenistan Tadzhikistan
Aphodius (Erytus) klugi A.Schmidt, 1910 Turkmenistan
Aphodius (Erytus) aequalis A.Schmidt, 1907 Turkmenistan
Aphodius (Erytus) bucharicus Petrovitz, 1961 Turkmenistan
Aphodius (Erytus) persicus Petrovitz, 1961 Afghanistan
Aphodius (Erytus) pruinosus Reitter, 1892 Turkmenistan
Aphodius (Plagiogonus) putridus (Fourcroy, 1785) RU UR WS SS Kazakhstan
Aphodius (Plagiogonus) praeustus Ballion, 1871 Kazakhstan Turkmenistan
Aphodius (Pharaphodius) rugosostriatus Waterhouse, 1875 Japan
Aphodius (Acrossus) bimaculatus (Laxmann, 1770) WS SS Kazakhstan
Aphodius (Acrossus) rufipes (Linnaeus, 1758) WS SS AM Italy Finland Ukraine Georgia Kazakhstan
Aphodius (Acrossus) depressus (Kugelann, 1798) RU CU UR WS SS Italy Ukraine Georgia
Aphodius (Acrossus) luridus (Fabricius, 1775) RU CU UR WS SS Crimea Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan Tadzhikistan
Aphodius (Acrossus) planicollis Reitter, 1890 Georgia
Aphodius (Acrossus) arsenjevi Berlov, 1989 AM
Aphodius (Aganocrossus) urostigma Harold, 1862 Japan
Aphodius (Neagolius) falcispinus W.Koshantschikov, 1912 SS
Aphodius (Neagolius) haroldi D.Koshantschikov, 1894 Kazakhstan
Aphodius (Neagolius) dzhungaricus Nikolajev et Frolov, 1997 paratypes! Kazakhstan
Aphodius (Gonaphodius) scoparius Harold, 1877 SS
Aphodius (Biralus) satellitus (Herbst, 1789) RU Kazakhstan
Aphodius (Biralus) menetriesi Menetries, 1849 Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan
Aphodius (Agolius) makowskyi D.Koshantschikov, 1891 Kazakhstan
Aphodius (Agolius) abchasikus Reitter, 1892 Georgia
Aphodius (Agoliinus) nemoralis Erichson, 1848 RU Belarus
Aphodius (Agoliinus) nasutus Reitter, 1887 SS
Aphodius (Agoliinus) tenebricosus A. Schmidt, 1816 WS SS
Aphodius (Aphodiellus) impunctatus Waterhouse, 1875 AM
Aphodius (Chilothorax) comma Reitter, 1892 WS SS MS Kazakhstan
Aphodius (Chilothorax) badenkoi Nikolajev, 1987 paratype! Kazakhstan
Aphodius (Chilothorax) flavimargo Reitter, 1901 Kazakhstan Turkmenistan
Aphodius (Chilothorax) distinctus (Muller, 1776) RU CU UR WS SS MS Kazakhstan Turkmenistan Tadzhikistan
Aphodius (Chilothorax) transvolgensis Semenov, 1898 RU WS Kazakhstan
Aphodius (Chilothorax) praenubilis Balthasar, 1938 Kyrghyzstan
Aphodius (Chilothorax) melanostictus W.Schmidt, 1840 RU WS MS Ukraine Kazakhstan Kyrghyzstan Turkmenistan
Aphodius (Chilothorax) auliensis Balthasar, 1933 Kyrghyzstan
Aphodius (Chilothorax) clathratus Reitter, 1892 unknown locality
Aphodius (Chilothorax) scuticollis Semenov, 1898 Kazakhstan Tadzhikistan
Aphodius (Chilothorax) sticticus (Panzer, 1798) RU CU Belarus
Aphodius (Chilothorax) grafi Reitter, 1901 SS
Aphodius (Chilothorax) tanhensis Frolov, 2001 SS
Aphodius (Chilothorax) jakobsoni W.Koshantshikov, 1911 SS Kazakhstan
Aphodius (Chilothorax) altaicus Nikolajev, 1984 China
Aphodius (Chilothorax) exilimanus Kabakov, 1994 UR
Aphodius (Chilothorax) kerzhneri Nikolajev, 1984 WS SS Kazakhstan
Aphodius (Chilothorax) mongolaltaicus Nikolajev, 1984 SS Mongolia
Aphodius (Melinopterus) prodromus (Brahm, 1790) RU CU UR WS SS Ukraine Latvia Kazakhstan Turkmenistan
Aphodius (Melinopterus) punctatosulcatus Sturm, 1805 RU UR WS SS MS Kazakhstan
Aphodius (Melinopterus) sphacelatus (Panzer, 1798) RU
Aphodius (Melinopterus) circumcinctus W.Schmidt, 1846 RU UR WS
Aphodius (Melinopterus) consputus Creutzer, 1899 RU
Aphodius (Melinopterus) scuticollis Semenov, 1898 Kazakhstan Tadzhikistan
Aphodius (Melaphodius) caspicus Menetries, 1823 RU WS SS MS Ukraine Kazakhstan
Aphodius (Nimbus) turkestanicus Heyden, 1881 Kazakhstan Kyrghyzstan
Aphodius (Nobius) serotinus (Panzer, 1799) RU WS Kazakhstan
Aphodius (Nobius) circumductus Solsky, 1876 Uzbekistan
Aphodius (Nobius) korgaldzhensis Nikolajev, 1987 UR
Aphodius (Nobius) pustulifer Reitter, 1892 Uzbekistan
Aphodius (Phaeaphodius) rectus Motschulsky, 1866 UR WS SS MS AM KU Kazakhstan China
Aphodius (Phaeaphodius) dauricus Harold, 1863 RU WS SS MS Kazakhstan
Aphodius (Phaeaphodius) costalis Gebler, 1848 WS Kazakhstan
Aphodius (Amidorus) tomentosus (Muller, 1776) WS SS Kazakhstan
Aphodius (Amidorus) obscurus (Fabricius, 1792) Georgia
Aphodius (Amidorus) zangi A.Schmidt, 1906 RU WS SS Kazakhstan
Aphodius (Orodaliscus) rotundangulus Reitter, 1900 RU WS Kazakhstan
Aphodius (Oromus) alpinus (Scopoli, 1763) Ukraine
Aphodius (Oromus) asphaltinus Kolenati, 1846 Georgia
Aphodius (Oromus) corvinus Erichson,  Belarus
Aphodius (Limarus) zenkeri (Germar, 1813) Belarus
Aphodius (Pseudacrossus) edgardi Solsky, 1876 Turkmenistan Tadzhikistan
Aphodius (Pseudacrossus) hahni Reitter,1907 UR Kazakhstan
Aphodius (Pseudacrossus) nasutus Reitter,  1887 SS
Aphodius (Pseudacrossus) grombczewskyi D.Koshantschikov, 1891 Kazakhstan Kyrghyzstan
Aphodius (Trichonotulus) scrofa (Fabricius, 1787)
Aphodius (Trichonotulus) scrofa scrofa (Fabricius, 1787) Kazakhstan
Aphodius (Trichonotulus) scrofa mongolicus (Mannerhaim, 1852) SS
Aphodius (Esymus) merdarius (Fabricius,  1775) RU CU Turkmenistan
Aphodius (Esymus) alaiensis Reitter, 1892 Kazakhstan Turkmenistan Tadzhikistan
Aphodius (Esymus) stereotypus D.Koshantschikov, 1894 Turkmenistan
Aphodius (Mecynodes) kisilkumi Solsky, 1876 WS Turkmenistan
Aphodius (Orodalus) pusillus (Herbst, 1789) RU CU UR WS SS MS AM KU Georgia Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan
Aphodius (Orodalus) coenosus (Panzer, 1798) RU UR Belarus
Aphodius (Eudolus) quadriguttatus (Herbst, 1783) UR Crimea Turkmenistan Uzbekistan
Aphodius (Phalacronotus) quadrimaculatus (Linnaeus, 1767) Turkmenistan
Aphodius (Phalacronotus) biguttatus Germar, 1824 UR WS Kazakhstan
Aphodius (Phalacronotus) citellorum Semenov et Medvedev, 1928 RU WS Kazakhstan
Aphodius (Mendidius) bidens Solsky, 1876 (Kazakhstan?) Turkmenistan
Aphodius (Mendidius) sijasovi Lebedev, 1932 Kazakhstan
Aphodius (Mendidius) multiplex Reitter, 1897 WS Kazakhstan Turkmenistan Mongolia
Aphodius (Mendidius) nelsinae (Medvedev, 1968) Kazakhstan
Aphodius (Mendidius) repetekensis (Nikritin, 1971) Turkmenistan
Aphodius (Mendidius) ivanovi Lebedev, 1912 RU
Aphodius (Mendidaphodius) brancsiki Reitter, 1899 Turkmenistan
Aphodius (Apsteiniella) georgii (W.Koshantschikov, 1912) Turkmenistan
Aphodius (Apsteiniella) aralicus Nikolajev, 1979 paratype! Kazakhstan
Aphodius (Loraspis) frater Mulsant et Ray, 1870 UR WS SS Kazakhstan
Aphodius (Loraphodius) suarius Faldermann, 1835 Azerbaijan
Aphodius (Labarrus) mathhiesseni Reitter, 1907 Afghanistan
Aphodius (Aphodius) fimetarius (Linnaeus, 1758) RU CU UR WS SS Ukraine Armenia Georgia Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan
Aphodius (Aphodius) foetens (Fabricius, 1787) RU UR WS SS Kazakhstan
Aphodius (Aphodius) elegans Allibert, 1848 Japan
Aphodius (Agrilinus) ater (De Geer, 1774) RU UR WS SS SH Georgia Kazakhstan
Aphodius (Agrilinus) borealis Gyllenhal, 1827 WS AM Finland Kazakhstan
Aphodius (Agrilinus) vittatus Say, 1825
Aphodius (Agrilinus) vittatus mundus Reitter, 1893 UR SS Kazakhstan Turkmenistan
Aphodius (Agrilinus) vittatus sellatus Mannerheim, 1852 SS
Aphodius (Agrilinus) uliginosus (Hardy, 1847) RU WS SS Belarus Kazakhstan
Aphodius (Agrilinus) lapponum Gyllenhal, 1806 SS Kazakhstan
Aphodius (Agrilinus) piceus Gyllenhal, 1808 RU SS Kazakhstan
Aphodius (Agrilinus) uniformis Waterhouse, 1875 KU
Aphodius (Agrilinus) pratensis Nomura et Nakane KU
Aphodius (Agrilinus) isajevi Kabakov, 1994 RU UR
Aphodius (Nialus) sturmi Harold, 1870 RU WS Kazakhstan Turkmenistan
Aphodius (Nialus) kraatzi Harold, 1868 Kazakhstan Kyrghyzstan Turkmenistan
Aphodius (Nialus) plagiatus (Linnaeus, 1767) WS SS Latvia Kazakhstan
Aphodius (Nialus) varians Duftschmid, 1805 RU WS Ukraine Azerbaijan Kazakhstan Tadzhikistan
Aphodius (Calamosternus) lividus (Olivier, 1789) AM Georgia Kazakhstan Turkmenistan Tadzhikistan
Aphodius (Calamosternus) granarius (Linnaeus, 1767) RU CU WS SS Georgia Azerbaijan Kazakhstan Turkmenistan Uzbekistan Tadzhikistan
Aphodius (Calamosternus) suturifer Reitter, 1893 SS Kazakhstan
Aphodius (Calamosternus) sublimbatus Motschulsky, 1860 AM
Aphodius (Aphodaulacus) koltzei Reitter, 1892 AM
Aphodius (Aphodaulacus) variabilis Waterhoze, 1875 AM

Melolonthinae
Melolonthini
Polyphylla (Mesopolyphylla) irrorata (Gebler, 1841) Kazakhstan IMAGES IN NATURE: pict.1
Polyphylla (Mesopolyphylla) tridentata Reitter, 1890 Kyrghyzstan
Polyphylla (Xerasiobia) alba (Pallas, 1773) CU WS Kazakhstan Turkmenistan
Polyphylla (Xerasiobia) adspersa Motschulsky, 1853 Turkmenistan Uzbekistan Tadzhikistan Kazakhstan
Polyphylla (Polyphylla) fullo (Linnaeus, 1758) RU Ukraine
Polyphylla (Polyphylla) olivieri (Laporte de Castelnau, 1840) CU Georgia Armenia
Polyphylla (Gynexophylla) laticollis Lewis, 1887 Japan
Lepidiota sp. Vietnam
Haplosternus incanus Motschulsky, 1853 AM
Melolontha (Melolontha) melolontha Linnaeus, 1758 Latvia
Melolontha (Melolontha) hippocostani Fabricius, 1801 WS SS
Melolontha (Melolontha) pectoralis Germar, 1824 CU Georgia
Melolontha (Apropyga) clypeata Reitter, 1887 Uzbekistan
Melolontha (Apropyga) afficta Ballion, 1870 Uzbekistan
Anoxia pilosa (Fabricius, 1792) RU

Asactopholis biturberculatus Moser, Vietnam

Heptophyllini
Heptophylla picea Motschulsky, 1857 South Korea Japan
Hilyotrogus bicoloreus Heyden, 1887 AM
Toxospathius auriventris Bates, 1891 China

Rhizotroginae
Rhizotrogini
Apogonia nigroolivaceae Heyden, 1886 AM
Apogonia sp. Rwanda
Lasiopsis (Lasiopsis) caninus Zoubkov, 1829 WS SS Kazakhstan
Lasiopsis (Lasiopsis) golovjankoi Medvedev, 1951 SS AM
Lasiopsis (Lasiopsis) sahlbergi Mannerhaim, 1849 SS
Lasiopsis (Brahmina) faldermanni Kraatz, 1892 SS
Lasiopsis (Brahmina) agnellus (Faldermann, 1835) SS
Lasiopsis (Brahmina) sedakovi (Mannerheim, 1849) SS AM
Lasiopsis (Brahmina) duchoni Reitter, 1902 Kazakhstan
Lasiopsis (Lachnota) henningi (Fischer, 1889) SS
Lasiopsis (Lasiexis) primaeveris (Semenov et Medvedev, 1936) Turkmenistan
Lasiopsis (Lasiexis) dilaticollis (Ballion, 1870) Kazakhstan Kyrghyzstan Turkmenistan
Cyphochilus insulanus Moser Taiwan
Lachnosterna sp. Rwanda
Panotrogus myschenkovi (Ballion, 1871) Kyrghyzstan Turkmenistan Tadzhikistan
Rhizotrogus (Rhizotrogus) aestivus (Olivier, 1789) RU Crimea Turkmenistan
Rhizotrogus (Amphimallon) altaicus (Mannerheim, 1825) WS SS Kazakhstan
Rhizotrogus (Amphimallon) volgensis (Fischer, 1823) Kazakhstan
Rhizotrogus (Amphimallon) solsticialis (Linnaeus, 1758)
Rhizotrogus (Amphimallon) solsticialis solsticialis (Linnaeus, 1758) Latvia
Rhizotrogus (Amphimallon) solsticialis setosus Reitter, 1902 Turkmenistan
Rhizotrogus (Amphimallon) solsticialis mesasiaticus Medvedev, 1951 Kazakhstan Kyrghyzstan Uzbekistan Tadzhikistan
Rhizotrogus (Amphimallon) solsticialis grossatus Reitter, 1902 Crimea
Rhizotrogus (Amphimallon) solsticialis sibiricus Reitter, 1902 WS SS
Rhizotrogus (Amphimallon) irtishensis Nikolajev, 1979 paratypes! Kazakhstan
Rhizotrogus (Amphimallon) ruficornis (Fabricius, 1775) SS
Xanthotrogus sieversi Reitter, 1896 Turkmenistan
Xanthotrogus fortis Reitter, 1902 Uzbekistan
Holochelus fallax (Marseul, 1879) Turkmenistan
Holochelus aequinoctialis (Herbst, 1790) RU Ukraine Kazakhstan
Holochelus tauricus (Blanchard, 1850) Ukraine
Holochelus (Madotrogus) glabripennis (Ballion, 1871) Kyrghyzstan Turkmenistan Uzbekistan Tadzhikistan
Holochelus (Dasitrogus) transcaspicus Brenske, 1866 Uzbekistan
Holochelus (Dasitrogus) glabricollis (Reitter, 1888) Turkmenistan
Holochelus (Dasitrogus) kirgisicus (Semenov, 1895) Kyrghyzstan
Holochelus (Dasitrogus) kuhitangicus Nikolajev, 1976 Turkmenistan
Chioneosoma (Aleucolomus) porosum (Fischer, 1824) Kazakhstan Turkmenistan
Chioneosoma (Aleucolomus) tedschenense Semenov, 1895 Turkmenistan
Chioneosoma (Aleucolomus) vulpinum (Gyllenhal, 1817) RU
Chioneosoma (Chionotrogus) candidum (Semenov, 1902) Turkmenistan
Chioneosoma (Chionotrogus) gorilla Brenske, 1886 Turkmenistan
Chioneosoma (Chionotrogus) kizilkumenze Semenov, 1895 Kazakhstan Turkmenistan
Chioneosoma (Chioneosoma) komarovi (Brenske, 1866) Turkmenistan
Chioneosoma (Chionotrogus) peetzi Semenov, 1898 Turkmenistan
Chioneosoma (Chionotrogus) pulvereum (Koch, 1801) RU
Chioneosoma (Chionotrogus) turcomanum Brenske, 1888 Turkmenistan
Chioneosoma (Leucolasium) baadeni Brenske, 1886 Turkmenistan
Chioneosoma (Leucolasium) rostovtzovi Semenov, 1898 Turkmenistan
Chioneosoma (Leucolasium) tschitscherini Semenov, 1895 Turkmenistan
Sophrops heydeni Brenske, 1892 AM
Sophrops planicollis Burmeister Vietnam
Sophrops cephalotes Burmeister Vietnam
Pectinichelus (Cryphaeobius) brunneus (Kraatz, 1882) Tadzhikistan
Pectinichelus (Cryphaeobius) lopatini Medvedev, 1959 Tadzhikistan
Holotrichia sichotana (Brenske, 1896) AM
Holotrichia diomphalia (Batten, 1895) AM China
Holotrichia inelegans (Lewis, 1895) Japan
Holotrichia lata Brenske China(Taiwan)
Holotrichia maxima Chang China
Holotrichia pruinosa Wied. Vietnam
Eotrichia titanis (Reitter, 1902) AM
Haplidia transversa (Fabricius, 1801) Bulgaria
Trematodes tenebrioides (Pallas, 1781) SS

Pachydeminae
Tanyproctus turanicus Reitter, 1900 Turkmenistan
Tanyproctus bucharicus Reitter, 1897 Tadzhikistan
Asiactenius tekkensis (Reitter, 1888) Turkmenistan
Asiactenius tokgajevi Medvedev, 1962 Turkmenistan
Hemictenius simplicitarsis Reitter, 1897 Tadzhikistan
Hemictenius ochripennis Reitter, 1902 Tadzhikistan
Hemictenius comatus Nikolajev, 1975 Tadzhikistan
Hemictenius lebedevi Reitter, 1908 Kazakhstan
Hemictenius lopatini Medvedev, 1959 Tadzhikistan
Hemictenius elongatus Medvedev, 1962 Tadzhikistan
Hemictenius pygmaeus Medvedev, 1962 Tadzhikistan
Hemictenius medvedevi Nikolajev, 1976 Turkmenistan
Hemictenius badkhyzicus Nikolaev, 2000 paratypes! Turkmenistan
Hemictenius opacus (Ballion, 1870) Uzbekistan
Hemictenius latitarsis (Reitter, 1891) Kazakhstan Turkmenistan Uzbekistan
Hemictenius magnitarsis (Reitter, 1897) Uzbekistan
Hemictenius nigrociliatus Reitter, 1897 Tadzhikistan
Hemictenius bactrianus Medvedev, 1952 Tadzhikistan
Hemictenius kryzhanovskii Nikolaev et Shukronajev, 1977 paratype! Tadzhikistan

Sericinae
Homaloplia marginata (Fuessly, 1775) Bulgaria
Homaloplia kiritshenkoi Medvedev, 1952 Crimea
Homaloplia hirta (Gebler, 1830) RU WS SS Kazakhstan
Homaloplia spiraea (Pallas, 1773) RU UR WS Armenia Kazakhstan
Homaloplia nicolasi Baraud, 1965 Italy
Serica (Serica) brunnea (Linnaeus, 1758) RU UR WS SS Ukraine Kazakhstan
Serica (Serica) ussuriensis Medvedev, 1952 AM
Serica (Serica) koltzei Reitter, 1897 AM
Serica (Ophthalmoserica) rosina Pic, 1904 AM
Serica (Ophthalmoserica) niijimai Konthwan Japan
Serica (Ophthalmoserica) karafutoensis Niijma et Kinoshita SH KU
Serica (Trichoserica) polita (Gebler, 1832) SS MS AM KU
Serica (Trichoserica) pilosa (Nomura) Japan
Maladera (Maladera) holosericea (Scopoli, 1772) RU CU SS Kazakhstan
Maladera (Maladera) renardi (Ballion, 1870) SS AM
Maladera (Aserica) punctatissima (Faldermann, 1835) Moldova Turkmenistan
Maladera (Aserica) euphorbia (Burmeister, 1895) Kazakhstan Turkmenistan Tadzhikistan
Maladera (Aserica) lukjanovitschi (Medvedev, 1966) SS
Maladera (Aserica) orientalis (Motschulsky, 1857) AM
Maladera (Aserica) castanea Arrow, 1913 SS AM Georgia
Maladera (Aserica) matrida Argamon, 1986 Israel
Maladera (Aserica) spissigrada (Brenske, 1897) AM
Leucoserica arenicola (Solsky, 1876) Turkmenistan
Sericania fuscolineata Motschulsky, 1860 AM SH
Sericania sachalinensis Matsumura, 1911 SH KU
Triodonta aquila (Castelnau, 1840) France

Hopliinae
Ectinohoplia rufipes (Motschulsky, 1860) AM SH KU
Ectinohoplia obducta (Motschulsky, 1857) Japan Taiwan
Ectinohoplia variabilis Reitter, 1903 Taiwan
Hoplia (Decamera) cincticollis (Faldermann, 1833) SS Mongolia
Hoplia (Decamera) hauseri Reitter, 1904 Kazakhstan Kyrghyzstan
Hoplia (Decamera) djukini Jacobson, 1914 AM
Hoplia (Hoplia) parvula Krynicky, 1832 RU
Hoplia (Hoplia) zaitzevi Jacobson, 1914 RU
Hoplia (Hoplia) detrita Solsky, 1876 RU
Hoplia (Hoplia) farinosa Linnaeus, 1761 Bavaria
Hoplia (Hoplia) graminicola Fabricius, 1792 Georgia
Hoplia (Hoplia) brevipes Medvedev, 1951 CU
Hoplia (Hoplia) pallinosa Krynicky, 1832 AM Georgia
Hoplia (Hoplia) praticola Duftschmidt, 1805 Bulgaria
Hoplia (Euchromoplia) aureola (Pallas, 1781) SS
Hoplia (Euchromorplia) communis Waterhouse, 1875 Japan
Hoplia ?elegantula Whit. Japan
Hoplia sp. China(Taiwan)

Cetoniinae
Trichiini
Gnorimus subopacus Motschulsky, 1860 SS AM
Gnorimus viridiopacus (Lewis, 1887) KU
Gnorimus octopunctatus Fabricius, 1775 RU
Gnorimus miwai (Chujo, 1941) China(Taiwan)
Lasiotrichius succinctus (Pallas, 1781) SS AM China
Trichius orientalis Reitter, 1894 Crimea
Trichius fasciatus (Linnaeus, 1787) RU CU UR WS SS AM SH KM Finland Latvia Ukraine Kazakhstan Finland
Osmoderma barnabita Motschulsky, 1860 AM
Osmoderma eremita Scopoly, 1763 RU

Goliathini
Rhomborrhina japonica (Hope, 1841) Japan
Rhomborrhina unicolor Motschulsky, 1861 Japan
Rhomborrhina polita Waterhoose, 1875 Japan
Trigonophorus rothschildi Fairmaire, 1891
Trigonophorus rothschildi varians (Bourgoin, 1914) China(Taiwan)
Stephanorrhina sp. Rwanda
Eudicella smithi (MacLeay, 1838) Rwanda
Diplognatha gagates (Forster, 1771) Rwanda

Gymnetini
Cotinis nitida (Linnaeus, 1764) USA
Coelodera penicillata (Hope, 1831)
Coelodera penicillata formosana China(Taiwan)
Euselatos proxima China(Taiwan)
Clynteria ducalis davidis Fairmaere China(Taiwan)
Clynteria 14-maculata (F.) Sri Lanka

Cetoniini
Epicometis (Epicometis) hirta (Poda, 1761) RU CU Crimea
Epicometis (Epicometis) hirtiformis Reitter, 1913 Kazakhstan Kyrghyzstan
Epicometis (Epicometis) spinifrons (Reitter, 1889) Turkmenistan
Epicometis (Epicometis) turanica (Reitter, 1888) Turkmenistan
Epicometis (Epicometis) suturalis Reitter, 1913 Armenia
Epicometis (Epicometis) senicula (Menetries, 1832) Armenia
Heterocnemis graeca (Brulle, 1832) Greece
Oxythyrea funesta (Poda, 1761) RU CU UR WS SS Crimea Kazakhstan Bulgaria
Oxythyrea albopicta (Motschulsky, 1845) Uzbekistan
Oxythyrea cinctella (Schaum, 1841) Armenia Kyrghyzstan Kazakhstan Turkmenistan UzbekistanTadzhikistan
Stalagmopygus albella (Pallas, 1781) Turkmenistan Tadzhikistan Afghanistan
Aethiessa szekessyi Brasavola, 1939 Tadzhikistan
Aethiessa mesopotamica Burmeister, 1842
Aethiessa mesopotamica rugipennis Burmeister, 1842 Turkmenistan
Cetonia (Eucetonia) magnifica Ballion, 1870 SS AM China
Cetonia (Eucetonia) viridiopaca (Motschulsky, 1860) SS CY AM
Cetonia (Eucetonia) roelofsi Harold, 1880 KU Japan
Cetonia (Cetonia) aurata (Linnaeus, 1761)
Cetonia (Cetonia) aurata aurata (Linnaeus, 1761) RU Latvia Belarus Ukraine Crimea Bulgaria Slovenia
Cetonia (Cetonia) aurata viridiventris Reitter, 1896 UR WS SS KU Kazakhstan Kyrghyzstan
Cetonia (Cetonia) aurata pisana Heer, 1891 Italy
Cetonia (Cetonia) aurata pallida (Drury, 1770) CU Armenia Georgia
Cetonia (Cetonia) aeratula Reitter, 1891 Azerbaijan
Protaetia (Liocola) lugubris (Herbst, 1786)
Protaetia (Liocola) lugubris lugubris (Herbst, 1786) RU WS SS Hungary Kazakhstan
Protaetia (Liocola) lugubris orientalis Medvedev, 1964 AM
Protaetia (Liocola) insperata (Lewis, 1879) KU
Protaetia (Calopotosia) mandschuriensis Smurhoff, 1933 AM
Protaetia (Calopotosia) brevitarsis (Lewis, 1879) AM
Protaetia (Cetonischema) aeruginosa (Drury, 1770) RU
Protaetia (Eupotosia) affinis (Andersch, 1797) RU
Protaetia (Potosia) cuprea (Fabricius, 1775)
Protaetia (Potosia) cuprea cuprea (Fabricius, 1775) France Italy
Protaetia (Potosia) cuprea obscura (Andersch, 1797) Bulgaria
Protaetia (Potosia) metallica (Herbst, 1782)
Protaetia (Potosia) metallica metallica (Herbst, 1782) RU UR WS SS Finland Latvia Ukraine Kazakhstan
Protaetia (Potosia) metallica daurica (Motschulsky, 1860) SS AM
Protaetia (Potosia) cuprina (Motschulsky, 1849) CU Crimea
Protaetia (Potosia) hieroglyphica (Menetries, 1832) Azerbaijan Turkmenistan
Protaetia (Potosia) marginicollis (Ballion, 1870) Kazakhstan Turkmenistan Uzbekistan Tadzhikistan
Protaetia (Potosia) angustata (Germar, 1817) Croatia
Protaetia (Potosia) lenzi (Harold) Japan
Protaetia (Netocia) funebris (Gory et Percheron, 1833) Turkmenistan
Protaetia (Netocia) turkestanica (Kraatz, 1886)
Protaetia (Netocia) turkestanica turkestanica (Kraatz, 1886) Turkmenistan
Protaetia (Netocia) turkestanica cyanea (Kraatz, 1886) Turkmenistan Uzbekistan Tadzhikistan
Protaetia (Netocia) bogdanovi (Solsky, 1875) Kyrghyzstan Tadzhikistan
Protaetia (Netocia) trojana (Gory et Percheron, 1833) Armenia Georgia
Protaetia (Netocia) prototricha (Fischer, 1842) Kazakhstan
Protaetia (Netocia) hungarica (Herbst, 1790)
Protaetia (Netocia) hungarica hungarica (Herbst, 1790) CU WS Latvia
Protaetia (Netocia) hungarica armeniaca (Menetries, 1832) Armenia
Protaetia (Netocia) hungarica pseudoviridana Medvedev, 1947 Turkmenistan
Protaetia (Netocia) hungarica inderiensis (Krynicky, 1832) Kazakhstan
Protaetia (Netocia) hungarica sibirica (Gebler, 1829) SS Mongolia
Protaetia (Netocia) persica (Kraatz, 1886) Turkmenistan
Protaetia (Netocia) excavata (Gory et Percheron, 1833) Turkmenistan
Protaetia (Netocia) interruptocostata (Ballion, 1870) Kyrghyzstan Tadzhikistan
Protaetia (Netocia) karelini (Zoubkov, 1829)
Protaetia (Netocia) karelini karelini (Zoubkov, 1829) UR Kazakhstan
Protaetia (Netocia) karelini agglomerata (Solsky, 1876) Turkmenistan Uzbekistan
Protaetia (Netocia) morio (Fabricius, 1781) Italy
Protaetia (Netocia) vidua (Gory & Percheron, 1833) Bulgaria
Protaetia (Netocia) asiatica (Faldermann, 1835) Azerbaijan
Protaetia (Pseudonetocia) kulabensis Reitter, 1893 Tadzhikistan
Protaetia (Protaetia) orientalis (Gory & Percheron, 1833) Japan
Protaetia culta Waterhouse 1879 Japan
Bricoptis variolosa (Gory & Percheron, 1833) Madagascar
Poecilophilides rusticola (Burmeister, 1842) AM Japan
Glycyphana fulvistemma (Motschulsky, 1860) AM China(Taiwan)
Glycyphana nepalensis
Glycyphana nepalensis roguesi Bourgoin Vietnam
Glycyphana georgievici Miksic Philippines
Glycyphana incongrua Janson, 1878 China(Taiwan)
Glycyphana pygmaea Mohnike Malaysia
Glycyphana cuculus Burmeister, 1842 Philippines
Oxycetonia jucunda (Faldermann, 1835) AM China
Oxycetonia forticula (Janson) Japan
Oxycetonia histrio (Olivier) Thailand
Pachnoda sinuata (Fabricius, 1775) Rwanda
Pachnoda peregrina marginata (Drury, 1773) Uganda laboratory

???ini
Anochilia nitida Pouillaude, 1915 Madagascar

Cremastochilini
Campsiura trguttulata Mohnike Indonesia
Clinterocera scabrosa Motschulsky, 1853 China

Valgini
Valgus hemipterus (Linnaeus, 1758) RU CU Crimea Kyrghyzstan
Dasyvalgus angusticollis (Waterhose, 1875) Japan
Dasyvalgus montivagus Moser Vietnam
Idiovalgus curtus Paulian, 1961 Vietnam
Idiovalgus ater Pic Vietnam

Dynastinae
Dynastini
Allomyrina dichotomus (Linnaeus, 1758) Japan

Phileurini
Eophileurus chinensis (Faldermann, 1835) Japan

Oryctini
Oryctes (Oryctes) nasicornis (Linnaeus, 1758)
Oryctes (Oryctes) nasicornis nasicornis (Linnaeus, 1758) RU CU WS SS Ukraine Crimea Armenia Kazakhstan Italy
Oryctes (Oryctes) nasicornis punctipennis Motschulsky, 1860 Tadzhikistan
Oryctes (Eremoryctes) ata Semenov et Medvedev, 1932 Turkmenistan
Oryctes (Howoryctes) sjostedti Kolbe Rwanda
Orizabrus clunalis (LeConte, 1856) USA
Strategus aloeus (Linnaeus, 1758) Cuba
Pentodon idiota (Herbst, 1798) CU Ukraine Crimea Iraq
Pentodon minutum Reitter, 1887 Turkmenistan
Pentodon bidens (Pallas, 1771)
Pentodon bidens bidens (Pallas, 1771) Azerbaijan Kazakhstan Turkmenistan Uzbekistan Tadzhikistan
Pentodon bidens punctatum Villers, 1899 Italy
Pentodon algerinum
Pentodon algerinum bispinifrons Reitter, 1894 Tadzhikistan
Pentodon quadridens (Gebler, 1845)
Pentodon quadridens quadridens (Gebler, 1845) Kazakhstan
Pentodon quadridens patruelis Frivaldszky, 1890 Mongolia
Pentodon sulcifrons Kuster, 1848 CU Azerbaijan
Podalgus cuniculus Burmeister, 1847
Podalgus cuniculus infantulus (Semenov, 1890) Turkmenistan
Phyllognathus excavatus (Forster, 1771) (=hauseri Reitter, 1894) Israel Turkmenistan Tadzhikistan
Eutyctus deserti Semenov, 1890 Turkmenistan
Pycnoschema diversum Peringuey, 1901 Rwanda

Xylotrupes gideon (Linnaeus)
Xylotrupes gideon australicus (Thomson) Australia

Rutelinae
Rutelini
Callistethus waterstraati ???
Callistethus waterstraati bawangensis Wada, 2000 paratype! Indonesia
Parastasia nigriceps Westwood, 1841 Indonesia
Peltonotus sulawesiensis Wada, 1990 paratype! Indonesia

Anomalini
Pharaonus lederi Reitter, 1888 Turkmenistan
Pharaonus semenovi Reitter, 1887 Turkmenistan
Phyllopertha diversa Waterhouse, 1875 Japan
Phyllopertha horticola (Linnaeus, 1758) RU CU WS SS AM Italy Kazakhstan China
Phyllopertha irregularis Waterhouse, 1875 Japan
Phyllopertha intermixta (Arrow, 1913) KU Japan
Phyllopertha arinicollis Ohaus, 1905 Vietnam
Phyllopertha ?humeralis Fairmaire China
Pseudosinghala dalmanni (Gyllenhal, ) Vietnam
Pseudosinghala transversa Burmaister Vietnam
Rhombonyx holosericea (Fabricius, 1787) RU WS SS SH
Rhombonyx pekinensis Heyden, 1886 China
Rhombonyx testaceipes (Motschulsky, 1860) Japan
Rhombonyx ussuriensis Medvedev, 1951 AM
Popillia atrocoerulea Bates, 1888 AM
Popillia indigonacea Motschulsky, 1853 AM China South Korea
Popillia quadriguttata (Fabricius, 1787)
Popillia quadriguttata quadriguttata (Fabricius, 1787) AM
Popillia quadriguttata japonica (Newmann, 1838) KU USA Japan
Popillia cyanea (Hope, 1831) Vietnam
Proagopertha lucidula (Faldermann, 1835) AM
Mimela flavilabris Waterhose, 1875 KU Japan
Mimela inonea Nomura Japan
Anomala (Euchronomala) cuprea (Hope, 1839) KU
Anomala (Euchronomala) viridana Kolbe, 1886 AM
Anomala (Euchronomala) mongolica Faldermann, 1835 AM
Anomala (Euchronomala) albopilosa Hope, 1839 Japan
Anomala edentula Ohaus Japan
Anomala (Anomala) vitis (Fabricius, 1775) Hungary
Anomala (Anomala) dubia (Scopoli, 1763) RU CU Latvia
Anomala (Anomala) abchasica Motschulsky, 1853 CU Georgia
Anomala (Anomala) lucens Ballion, 1871 SS KU
Anomala (Anomala) daimiana Harold, 1877 Japan
Anomala (Anomala) rufocuprea Motschulsky, 1860 KU Japan
Anomala (Anomala) motschulskyi Harold, 1877 Japan
Anomala (Anomala) ogloblini Medvedev, 1951 AM
Anomala (Anomala) luculenta Erichson, 1847 SS AM
Anomala (Anomala) cupripes Hope Vietnam
Anomala (Anomala) antiqua Gyllenhal, 1817 Vietnam
Anomala (Psammoscaphaeus) vittata Gebler, 1841 Kazakhstan
Anomala (Psammoscaphaeus) metonidia Reitter, 1903 Kyrghyzstan
Anomala (Psammoscaphaeus) errans (Fabricius, 1775) RU Crimea Ukraine
Anomala (Psammoscaphaeus) oxiana Semenov, 1891 Tadzhikistan
Anomala dimidiata (Hope, 1831) Afghanistan
Anomala barbata Burmeister Afghanistan
Anomala costata Hope Japan
Trigonocnemis (Ammogenia) varentzovi Semenov, 1895 Turkmenistan
Blitopertha (Exomala) pallidipennis Reitter, 1903 AM
Blitopertha (Exomala) orientalis Waterhose, 1875 Japan
Blitopertha (Blitopertha) variabilis Ballion, 1870 Kazakhstan Turkmenistan Uzbekistan Tadzhikistan
Blitopertha (Blitopertha) lineata Fabricius, 1798 RU Armenia Azerbaijan Turkmenistan
Blitopertha (Blitopertha) conspurcata Harold, 1878 AM
Blitopertha (Asiopertha) arenicola Mulsant, 1870 CU
Cyriopertha (Cyrioperta) glabra (Gebler, 1841) Kazakhstan Uzbekistan Tadzhikistan
Cyriopertha (Cyrioperta) arcuata (Gebler, 1832) SS
Cyriopertha (Megapertha) massageta Kirsch, 1881 Turkmenistan Tadzhikistan
Anisoplia (Chaetopteroplia) leucaspis Laporte, 1840 Turkmenistan
Anisoplia (Chaetopteroplia) segetum (Herbst, 1783)
Anisoplia (Chaetopteroplia) segetum segetum (Herbst, 1783) RU Bulgaria
Anisoplia (Chaetopteroplia) segetum zoubkovi Krynicky, 1832 Kazakhstan
Anisoplia (Chaetopteroplia) syriaca Burmeister, 1844 Crimea
Anisoplia (Autanisoplia) austriaca (Herbst, 1783) RU Ukraine
Anisoplia (Lasioplia) campicola Menetries, 1832 RU UR WS SS Kazakhstan
Anisoplia (Lasioplia) koenigi Reitter, 1895 Turkmenistan
Anisoplia (Lasioplia) signata Faldermann, 1835 Armenia Georgia
Anisoplia (Lasioplia) brenskei Reitter, 1889 RU Georgia
Anisoplia (Anisoplia) farraria Erichson, 1847 Armenia
Anisoplia (Anisoplia) turcomana Zaitzev, 1917 Turkmenistan
Anisoplia (Anisoplia) agricola (Poda, 1761) RU CU WS Kazakhstan
Anisoplia (Anisoplia) zwicki Fischer, 1823 RU
Anisoplia (Ammanisoplia) deserticola Fischer, 1823 RU WS
Parastasia nigriceps Westwood, 1841 Philippines

Adoretini
Phaeadoretus comptus (Menetries, 1849) Kazakhstan Turkmenistan Uzbekistan
Adoretus pruinosus Ballion, 1870 Kazakhstan
Adoretus nigrifrons (Steven, 1809) RU CU Turkmenistan Uzbekistan Kyrghyzstan Tadzhikistan
Adoretus discolor (Faldermann, 1835) Armenia
Adoretus rufifrons Reitter, 1903 Mongolia
Adoretus convexus Burmeister, Vietnam
Adoretus sinicus Burmeister, 1844 Vietnam
Adoretus (Lepadoretus) tenuimaculatus Waterhouse, 1875
Pseudoadoretus validus Semenov, 1889 Turkmenistan
Pseudoadoretus dilutellus Semenov, 1889 Turkmenistan
Pseudoadoretus phthisicus Dohrn, 1882 Uzbekistan