materials from different regions of Russia are encoded as: FS RU CU UR WS SS MS CY NE NF KM AM SH KU
MACROPYLINA
Paleosomata Grandjean, 1969
PALAEACAROIDEA Grandjean, 1932
PALAEACARIDAE Grandjean, 1932
Palaeacarus kamenskii (Zachvatkin, 1945) WS
SS MS KU
CTENACAROIDEA Grandjean, 1954
CTENACARIDAE Grandjean, 1954
Gilarovella demetrii Lange, 1974 RU Kazakhstan
APHELACARIDAE Grandjean, 1954
Aphelacarus acarinus (Berlese, 1910) RU
Arthronota Balogh, 1972
HAPLOCHTHONIOIDEA Van der Hammen, 1959
HAPLOCHTHONIIDAE Van der Hammen, 1959
Haplochthonius simplex (Willmann, 1930)
WS
Georgia
Haplochthonius sanctaeluciae Bernini, 1973 Turkmenistan
BRACHYCHTHONIOIDEA Thor, 1934
BRACHYCHTHONIIDAE Balogh, 1943
Brachychochthonius cricoides (Weis-Fogh, 1848 ) WS
SS Kazakhstan
Brachychochthonius hungaricus (Balogh, 1943) RU
Brachychochthonius immaculatus Forsslund, 1942 RU
WS Kazakhstan
Brachychochthonius jacoti (Ewans, 1952) WS
Brachychochthonius jugatus Jacot, 1938 Poland
Brachychochthonius meridionalis Bernini, 1973 RU
WS
Brachychochthonius rostratus (Jacot,1936)
WS SS
Brachychthonius berlesei Willmann, 1928 Kazakhstan
Brachychthonius bimaculatus Willmann, 1936
Poland
Brachychthonius hirtus Moritz, 1976 RU
Brachychthonius impressus Moritz, 1976 RU
Eobrachychthonius latior (Berlese, 1910) WS
Eobrachychthonius oudemansi Van der Hammen, 1952 Turkmenistan
Liochthonius alpestris (Forsslund, 1958) WS
Liochthonius evansi (Forsslund, 1952) MS
Liochthonius hystricinus (Forsslund, 1942) SS
MS
Liochthonius laetepictus (Berlese, 1910) MS
Liochthonius lapponicus (Trägårdh, 1910) RU
SS
Liochthonius leptaleus Moritz, 1976 RU
Liochthonius perpusillus (Berlese, 1910)WS
SS MS
Liochthonius propinquus Niedbala, 1972 Poland
Liochthonius sellnicki (Thor, 1930) WS
SS MS Kazakhstan
Liochthonius simplex (Forsslund, 1942) RU
Poecilochthonius italicus (Berlese, 1910) WS
SS
Synchthonius crenulatus (Jacot, 1938) SS
Synchthonius elegans Forsslund, 1957 WS
MS
HYPOCHTHONIOIDEA Berlese, 1910
HYPOCHTHONIIDAE Berlese, 1910
Hypochthonius luteus Oudemans, 1917 WS
Azerbaijan
Hypochthonius rufulus C.L. Koch, 1836 RU
WS SS AM KU Belarus Georgia
ENIOCHTHONIOIDEA Grandjean, 1947
ENIOCHTHONIIDAE Grandjean, 1947
Hypochthoniella minutissima (Berlese, 1904) WS
SS AM KU Belarus Moldova
COSMOCHTHONIOIDEA Grandjean, 1947
COSMOCHTHONIIDAE Grandjean, 1947
Cosmochthonius lanatus (Michael, 1885) Georgia
Cosmochthonius novus Sergienko, 1991 MS
Cosmochthonius plumatus Berlese, 1910 Uzbekistan
Cosmochthonius reticulatus (?) Grandjean, 1947 Turkmenistan
Phyllozetes tauricus Gordeeva, 1980 RU
WS
Israel
SPHAEROCHTHONIOIDEA Grandjean, 1947
SPHAEROCHTHONIIDAE Grandjean, 1947
Sphaerochthonius splendidus (Berlese, 1904)
Crimea Georgia Azerbaijan Tadzhikistan Kazakhstan
Sphaerochthonius ovatus Sergienko, 1991 Georgia
Sphaerochthonis sp. Turkmenistan
HETEROCHTHONIOIDEA Grandjean, 1954
HETEROCHTHONIIDAE Grandjean, 1954
Heterochthonius gibbus Berlese, 1910 WS
SS Georgia
ATOPOCHTHONIOIDEA Grandjean, 1948
ATOPOCHTHONIIDAE Grandjean, 1948
Atopochthonius maimaensis Grishina, 1971 paratype!
SS
MESOPLOPHOROIDEA Ewing, 1917
MESOPLOPHORIDAE Ewing, 1917
Mesoplophora pulchra Sellnick, 1928 Czechia
Parhyposomata Balogh et Mahunka, 1979
PARHYPOCHTHONIOIDEA Grandjean, 1932
PARHYPOCHTHONIIDAE Grandjean, 1932
Parhypochthonius aphidinus Berlese, 1904 RU
GEHYPOCHTHONIIDAE Strenzke, 1963
Gehypochthonius rhadamanthus Jacot, 1936 RU
Mixonomata Grandjean, 1969
LOHMANNIOIDEA Berlese, 1916
LOHMANNIIDAE Berlese, 1916
Mixacarus exilis Aoki, 1970 Azerbaijan
Papillacarus akimovi Sergienko, 1992 Kazakhstan
Papillacarus pavlovski (Bulanova-Zachvatkina, 1960) CU
Papillacarus sp. Azerbaijan
Thamnacarus longisetosus Bulanova-Zachvatkina, 1978
paratype! Turkmenistan Uzbekistan
Thamnacarus smirnovi Bulanova-Zachvatkina, 1978 paratype!
Turkmenistan
Vepracarus sp. RU
EULOHMANNIOIDEA Grandjean, 1931
EULOHMANNIIDAE Grandjean, 1931
Eulohmannia ribagai Berlese, 1910 SS SH
PERLOHMANNIOIDAE Grandjean, 1954
PERLOHMANNIIDAE Grandjean, 1954
Perlohmannia altaica Grishina, 1968 paratype!
SS
Perlohmannia coiffaiti Grandjean, 1961 KU
Perlohmannia zachvatkini (Bulanova-Zachvatkina, 1960) AM
EPILOHMANNIOIDEA Oudemans, 1923
EPILOHMANNIIDAE Oudemans, 1923
Epilohmannia cylindrica (Berlese, 1904) RU
WS SS Georgia Azerbaijan Kazakhstan
Epilohmannia spatulata Aoki, 1970 AM
Neolohmannia ovata (Aoki, 1961) AM
PHTHIRACAROIDEA Perty, 1841
PHTHIRACARIDAE Perty, 1841
Archiphthiracarus globus (Parry, 1979) SS
MS
Archiphthiracarus pilosus (Sergienko, 1987)
paratype! MS
Ukraine
Hoplophthiracarus candidulus Niedbala, 1983
"Europe"
Georgia
Hoplophthiracarus meridionalis Sergienko, 1982 Crimea
Hoplophthiracarus sp. Tajikistan
Phthiracarus assimilis Niedbala, 1983 Crimea
Georgia
Phthiracarus conformis Sergienko, 1987 paratype!
Ukraine
Phthiracarus crassus Niedbala, 1983 CU
Phthiracarus dissonus Niedbala, 1983 WS
SS MS
Phthiracarus globosus (C.L.Koch, 1841) RU
UR WS SS MS AM
Phthiracarus incredibilis Niedbala, 1983 WS
Phthiracarus italicus (Oudemans, 1900) Ukraine
Phthiracarus kaszabi Mahunka, 1981 WS
MS KU
Phthiracarus laevigatus (C.L.Koch, 1841) SS
Ukraine
Phthiracarus pallidus Feider et Suciu, 1958 Georgia
Phthiracarus propinquus Niedbala, 1983 "Europe"
Steganacarus anomalus (Berlese, 1883) CU
Georgia
Steganacarus brevipilus (Berlese, 1923) Ukraine
Azerbaijan
Steganacarus carinatus (C.L.Koch, 1841) Crimea
Steganacarus magnus (Nicolet, 1855) CU
Steganacarus ochraceus Niedbala, 1983 Ukraine
Georgia
Steganacarus patruelis Niedbala, 1983 CU
Georgia
Steganacarus personatus Niedbala, 1983 CU
Steganacarus pulcherrhimus (Berlese, 1887) "Europe"
Georgia Azerbaidjan
Steganacarus punctulatus Sergienko, 1985 paratype!
Ukraine
Steganacarus serratus Feider et Suciu, 1957 Ukraine
Steganacarus striculus (C.L.Koch, 1836) RU
WS SS AM Ukraine
EUPHTHIRACAROIDEA Jacot, 1930
ORIBOTRITIIDAE Grandjean, 1954
Maerkelotritia krivolutzkii Märkel, 1968 MS
Mesotritia nuda (Berlese, 1887) Ukraine
Mesotritia piffli Märkel, 1964 "Middle
Asia"
Mesotritia testacea Forsslund, 1963 RU
SS
Oribotritia hermannii Grandjean, 1967 Azerbaijan
Oribotritia kishidai Aoki, 1958 KU
Oribotritia tokukoae Aoki, 1973 KU
Oribotritia sp. Georgia
Protoribotritia oligotricha Märkel, 1963 WS
EUPHTHIRACARIDAE Jacot, 1930
Euphthiracarus cribrarius (Berlese, 1904) RU
WS SS MS
Euphthiracarus sp. MS
Microtritia fissurata Märkel, 1968 WS
SS MS
Microtritia minima (Berlese, 1904) Ukraine
Rhysotritia ardua affinis Sergienko, 1989 paratype!
UR
WS SS MS Ukraine Kazakhstan
Rhysotritia ardua ardua (C.L.Koch, 1841) RU
WS MS AM Israel
Rhysotritia ardua penicillata Perez-Inigo, 1969 MS
Tajikistan
Turkmenistan
Rhysotritia duplicata (Grandjean, 1953) Ukraine
Rhysotritia sp. AM
Holosomata Grandjean, 1969
NOTHROIDEA Berlese, 1885
NOTHRIDAE Berlese, 1896
Nothrus anauniensis Canestrini et Fanzago, 1878 Belarus
Kazakhstan
Nothrus biciliatus C.L. Koch, 1841 WS
SS MS AM Lithuania Georgia Kazakhstan
Nothrus borussicus Sellnick, 1928 RU WS
SS MS Kazakhstan
Nothrus palustris C.L. Koch, 1839 WS SS
MS AM
Nothrus parvus Sitnikova, 1975 WS MS
Nothrus pratensis Sellnick, 1928 WS SS
MS NF
Nothrus quadripilis Ewing, 1909 USA
Nothrus reticulatus Sitnikova, 1975 AM
KU
Nothrus silvestris Nicolet, 1855 WS Georgia
Azerbaijan
Nothrus terminalis Banks, 1910 USA
CAMISIIDAE Oudemans, 1900
Camisia biurus (C.L.Koch, 1839) SS
Camisia biverrucata (C.L.Koch, 1839) WS
SS NF "Europe" Kazakhstan
Camisia biverrucata copetdaghi Tolstikov, 1997 HOLOTYPE!
Turkmenistan
Camisia foveolata Hammer, 1955 MS
Camisia horrida (Hermann, 1804) CU UR
WS
"Europe" Lithuania
Camisia invenusta (Michael, 1888) UR
Camisia lapponica (Trägårdh, 1910) SS
Camisia segnis (Hermann, 1804) CU "Europe"
Middle Asia
Camisia solhoeyi Colloff, 1993 WS KU NE
Camisia spinifer (C.L.Koch, 1836) SS KU
Azerbaijan Kazakhstan
Heminothrus humicola (Forsslund, 1955) WS
SS
Heminothrus longisetosus (Willmann, 1925) SS
KU
Heminothrus peltifer (C.L. Koch, 1839) RU
WS SS Lithuania Ukraine Kazakhstan
Heminothrus punctatus (L. Koch, 1879) WS
Heminothrus septentrionalis Sellnick, 1944 KM
KU
Heminothrus sibiricus (Sitnikova, 1975) RU
WS MS
Heminothrus targionii (Berlese, 1885) CU
WS SS AM Crimea
Heminothrus thorii (Berlese, 1904) WS
Georgia
Heminothrus yamasakii (Aoki, 1958) AM
KU
TRHYPOCHTHONIIDAE Willmann, 1931
Hydronothrus crispus Aoki, 1964 KU
Mucronothrus nasalis (Willmann, 1929)
Georgia
Trhypochthoniellus setosus (Willmann, 1928) WS
Trhypochthonius cladonicola (Willmann, 1919) WS
NE
Trypochthonius septentrionalis Fujikawa, 1995 KU
Trhypochthonius setantis Golosova, 1983 SS
Georgia
Kazakhstan
Trhypochthonius tectorum (Berlese, 1896) WS
SS MS AM
MALACONOTHRIDAE Berlese, 1916
Malaconothrus mollisetosus Hammer, 1952 WS
AM
Malaconothrus punctulatus Van der Hammen, 1952 MS
Malaconothrus pygmaeus Aoki, 1969 WS AM
KU Kazakhstan Georgia
Trimalaconothrus glaber (Michael, 1888) WS
Trimalaconothrus novus (Sellnick, 1921) WS
Crimea
Kazakhstan
NANHERMANNIOIDEA Sellnick,1928
NANHERMANNIIDAE Sellnick, 1928
Nanhermannia comitalis Berlese, 1916 WS
Lithuania
Nanhermannia coronata Berese, 1913 RU
WS KU Lithuania
Nanhermannia dorsalis (Banks, 1896) USA
Nanhermannia elongatula Berlese, 1913 KU
Nanhermannia komareki Kunst, 1956 "Europe"
Nanhermannia laevis Sitnikova, 1975 KU
Nanhermannia nana (Nicolet, 1855) AM Crimea
Lithuania
Nanhermannia pectinata Strenzke, 1958 WS
SS
Nanhermannia sellnicki Forsslund, 1958 AM
HERMANNIOIDEA Sellnick, 1928
HERMANNIIDAE Sellnick, 1928
Hermannia gibba (C.L.Koch, 1840) "Europe"
Hermannia gigantea Sitnikova, 1975 WS
MS
Hermannia scabra ( C.L.Koch, 1879) MS
Hermannia sp. AM
BRACHYPYLINA
Apterogasterina-Gymnonota
HERMANNIELLOIDEA Grandjean, 1934
HERMANNIELLIDAE Grandjean, 1934
Hermanniella dolosa Grandjean, 1931 AM
LIODOIDEA Grandjean, 1954
LIODIDAE Grandjean, 1954
Liodes theleproctus (Hermann, 1804) CU
WS
Kazakhstan
Platyliodes scaliger (C.L.Koch, 1839) "Europe"
Georgia
Poroliodes farinosus (C.L.Koch, 1839) KU
GYMNODAMAEOIDEA Grandjean, 1954
GYMNODAMAEIDAE Grandjean, 1954
Gymnodamaeus bicostatus (C.L.Koch, 1835) WS
Kazakhstan
Nortonella mongolica Bayartogtokh, Aoki, 1997 MS
Kazakhstan
Nortonella (?) sp. MS AM
Jonstonella (?) sp. WS
Pleodamaeus kazakhstanicus Bayartogtokh et Smelyansky, 2002 HOLOTYPE! paratypes!
Kazakhstan
Pleodamaeus tuberculatus Bayartogtokh et Smelyansky, 2002 HOLOTYPE! paratypes!
RU
Plesiodamaeus glaber Mihelcic, 1835 RU
PLATEREMAEOIDEA Trägårdh, 1931
PLATEREMAEIDAE Trägårdh, 1931
Allodamaeus sp. "Europe"
PEDROCORTESELLIDAE Paschoal, 1987
Pedrocortesella fusca (Ryabinin, 1986) Kazakhstan
Pedrocortesella inaequalis (Balogh et Mahunka, 1965) SS
AM Kazakhstan
Pedrocortesella minuta Bayartogtokh et Smelyansky, 2004 HOLOTYPE! paratypes!
RU
Pedrocortesella rarisetosa Bayartogtokh et Smelyansky, 2004 HOLOTYPE! paratypes!
Kazakhstan
LICNODAMAEIDAE Grandjean, 1954
Licnodamaeus pulcherrimus (Paoli,1908)
RU WS "Europe"
Licnodamaeus fissurata (Balogh et Mahunka, 1965) AM
Kazakhstan
DAMAEOIDEA Berlese, 1896
DAMAEIDAE Berlese, 1896
Belba corynopus (Hermann,1804) CU Germany
Belba dubinini Bulanova-Zachvatkina, 1962 unknown
locality
Belba geniculosa Oudemans, 1929 Germany
Czechia
Belba meridionalis Bulanova-Zachvatkina, 1962 Turkmenistan
Belba paracorynopus Bulanova-Zachvatkina, 1962 AM
China
Belba rossica Bulanova-Zachvatkina, 1962 WS
SS
Belba sellnicki Bulanova-Zachvatkina, 1962 AM
Belba verrucosa Bulanova-Zachvatkina, 1962 WS
Caenobelba alleganiensis Norton, 1980 Canada
Damaeus auritus C.L.Koch,1835 KU
Damaeus gracilipes (Kulczynski, 1902) unknown
locality
Epidamaeus aborigensis Behan-Pelletier et Norton, 1985 paratype!
NF
Epidamaeus affinis Bulanova-Zachvatkina, 1957 paratype!
RU
WS MS
Epidamaeus aleinikovae Bulanova-Zachvatkina, 1964 paratype!
RU
Epidamaeus arcticolus (Hammer, 1952) USA(Alaska)
Epidamaeus bakeri (Hammer, 1952) USA(Alaska)
Epidamaeus bituberculatus (Kulczynski, 1902) WS
SS MS
Epidamaeus canadensis (Banks, 1909) Canada
Epidamaeus coxalis (Hammer, 1952) USA(Alaska)
Epidamaeus floccosus Behan-Pelletier et Norton, 1985 USA(Alaska)
Epidamaeus fortispinosus Hammer,1967 KU
Epidamaeus golosovae Ljastshev et Tolstikov, 1993 paratype!
Uzbekistan
Epidamaeus grandjeani Bulanova-Zachvatkina, 1957 AM
Epidamaeus hammerae Behan-Pelletier et Norton, 1983 paratype!
NF
Epidamaeus karelicus Bulanova-Zachvatkina, 1957 RU
Epidamaeus kodiakensis Hammer, 1967 USA(Alaska)
Epidamaeus koyukon Behan-Pelletier, 1985 Canada
Epidamaeus mackenziensis (Hammer, 1952) USA(Alaska)
Epidamaeus microspinus Bulanova-Zachvatkina, 1957 paratype!
AM
KU
Epidamaeus nasutus Behan-Pelletier et Norton, 1985 paratype!
USA(Alaska)
Epidamaeus paraspinosus Bulanova-Zachvatkina, 1974 MS
Epidamaeus plumosus Bulanova-Zachvatkina, 1974 paratype!
RU
Epidamaeus pseudotatricus Bulanova-Zachvatkina, 1957
RU UR
Epidamaeus tritylos Behan-Pelletier, 1983 USA(Alaska)
Hypodamaeus brevitibialis Bulanova-Zachvatkina, 1955 paratype!
RU
Hypodamaeus bulbofemoralis Bulanova-Zachvatkina, 1957 paratype!
Kyrghyzstan
Hypodamaeus crispatus (Kulczynski,1902) Crimea
Hypodamaeus echinopus Bulanova-Zachvatkina, 1957 paratype!
CU
Hypodamaeus riparius (Nicolet,1855) RU
Crimea
Hypodamaeus smirnovi Bulanova-Zachvatkina, 1957 Crimea
Hypodamaeus tauricus Bulanova-Zachvatkina, 1957 Crimea
Hypodamaeus tenuitibialis Bulanova-Zachvatkina, 1957
HOLOTYPE! paratype! Kyrghyzstan
Metabelba ericius Kunst, 1961 Ukraine
Metabelba filippovi Bulanova-Zachvatkina, 1965 CU
Georgia
Azerbaijan
Metabelba flagelliseta Bulanova-Zachvatkina, 1965 Crimea
Metabelba monilipeda Bulanova-Zachvatkina, 1967 paratype!
CU
Metabelba papillipes (Nicolet,1855) RU
Metabelba pseudoitalica Bulanova-Zachvatkina, 1965 CU
Metabelba pulverulenta (C.L.Koch, 1839)
RU CU Germany Hungaria
Metabelba rara Bulanova-Zachvatkina, 1965 paratype!
Georgia
Metabelba rohdendorfi Bulanova-Zachvatkina, 1967 RU
CU UR Czechia
Metabelbella langei Bulanova-Zachvatkina, 1960 CU
Metabelbella macerochaeta Bulanova-Zachvatkina, 1967 Georgia
Metabelbella tichonravovi Bulanova-Zachvatkina, 1965 Crimea
Metabelbella zachvatkini Bulanova-Zachvatkina, 1957 Azerbaijan
Parabelbella crenatosetosa (Bulanova-Zachvatkina, 1957) Kyrghyzstan
Parabelbella elisabethae Bulanova-Zachvatkina, 1967 WS
MS
Porobelba spinosa (Sellnick, 1920) WS
Spatiodamaeus boreus Bulanova-Zachvatkina, 1957 SS
Spatiodamaeus fageti Bulanova-Zachvatkina, 1957 unknown
locality
Spatiodamaeus subverticillipes Bulanova-Zachvatkina, 1957
SS
Spatiodamaeus verticillipes (Nicolet,1855) RU
UR Crimea
CEPHEOIDEA Berlese, 1896
CEPHEIDAE Berlese, 1896
Cepheus dentatus (Michael, 1888) unknown
locality
Cepheus cepheiformis (Nicolet, 1855) FS
MS
Tritegeus bifidatus (Nicolet, 1855) unknown
locality
Tritegeus brevisetus Sitnikova, 1975 SS
CHARASSOBATOIDEA Grandjean, 1958
CHARASSOBATIDAE Grandjean,
1958
Ametroproctus beringianus Behan-Pelletier, 1987 Canada
Ametroproctus canningsi Behan-Pelletier, 1987 Canada
Ametroproctus oresbios Higgins et Wooley, 1968 Canada
AMEROBELBOIDEA Grandjean, 1954
AMERIDAE Grandjean, 1965
Amerus troisii (Berlese, 1883) unknown
locality
DAMEOLIDAE Grandjean, 1965
Damaeolus asperatus (Berlese, 1904) unknown
locality
Fosseremus laciniatus (Berlese, 1905) SSKazakhstan
EREMAEOIDEA Woolley, 1965
EREMAEIDAE Sellnick,1928
Eremaeus hepaticus C.L.Koch, 1835 SS MS
Eremaeus insertus Grishina, 1980 paratype!
WS
MS
Eremaeus orientalis Golosova, 1970 AM
Proteremaeus elongatus (Krivolutsky et Rjabinin, 1975) SS
Proteremaeus jonasi Piffl, 1965 WS SS
Proteremaeus nebaikini Behan-Pelletier et Rjabinin, 1991 HOLOTYPE!
AM
ZETORCHESTOIDEA Baligh, 1961
ZETORCHESTIDAE Michael, 1898
Microzetorchestes emeryi (Coggi, 1898) Kazakhstan
Zetorchestes micronychus (Berlese, 1883) unknown
locality
GUSTAVIOIDEA Oudemans, 1900
ASTEGISTIDAE Balogh, 1961
Astegistes pilosus (C.L.Koch, 1840) WS
Cultroribula bicultrata (Berlese, 1905) WS
KU
Cultroribula lata Aoki, 1961 SS
Cultroribula ligulata Grishina, 1980 paratype!
MS
Cultroribula microdentata Krivolutsky, 1962 KU
Cultroribula tridentata Aoki, 1965 AM
Cultroribula vtorovi Krivolutsky, 1971 WS
SS MS
Furcoribula furcillata (Nordenskiold, 1901) Ukraine
GUSTAVIIDAE Oudemans, 1900
Gustavia microcephala (Nicolet, 1855) WS
SS
LIACARIDAE Sellnick, 1928
Adoristes poppei (Oudemans, 1906) WS
Birstenius microchaetus Krivolutsky, 1967 WS
SS MS
Birstenius perlongus Krivolutsky, 1965 WS
SS MS
Dorycranosus acutidens (Aoki, 1970) KU
Dorycranosus altaicus Krivolutsky, 1974 SS
Dorycranosus arcticus Grishina, 1984 paratype!
UR
WS SS
Liacarus cuspidatus Mihelcic, 1954 Georgia
Liacarus lencoranicus Krivolutsky, 1967 RU
CU WS MS
Liacarus xylariae (Schrank, 1803) WS SS
METRIOPPIDAE Balogh, 1943
Ceratoppia bipilis (Hermann, 1804) WS
SS MS
Ceratoppia quadridentata (Haaaller, 1882) WS
Ceratoppia sexpilosa Willmann, 1938 MS
Paraceratoppia meridionalis Rjbinin, 1982 KM
Pyroppia arctica Krivolutsky, 1974 WS
TENUIALIDAE Jacot,1929
Hafenrefferia gilvipes (C.L.Koch, 1839) unknown
locality
Peltenuiala pacifica Norton, 1983 Canada
XENILLIDAE Woolley et Higgins, 1966
Xenillus discrepans Grandjean, 1936 WS
SS
Xenillus stepensis Dzhaparidze, 1974 CU
Xenillus tegeocranus (Hermann, 1804) WS
MS KU Kazakhstan
CARABODOIDEA C.L.Koch, 1837
CARABODIDAE C.L.Koch, 1837
Carabodes areolatus Berlese, 1916 WS
Carabodes coriaceus C.L.Koch, 1835 Azerbaijan
Carabodes femoralis (Nicolet, 1855) FS
Belarus
Carabodes forsslundi Sellnick, 1953 Belarus
Carabodes labyrinthicus (Michael, 1879) SS
MS Belarus
Carabodes marginatus (Michael, 1884) Kenia
Carabodes minusculus Berlese, 1923 Belarus
Carabodes reticulatus Berlese, 1913 unknown
locality
Carabodes subarcticus Trägårdh, 1902 Azerbaijan
Kenia
Carabodes tenius Forsslund, 1953 unknown
locality
TECTOCEPHEIDAE Grandjean, 1954
Tectocepheus knullei Vanek, 1960 WS SS
Tectocepheus punctulatus Dzhaparidze, 1985 paratype!
Crimea
Georgia Azerbaijan
Tectocepheus sarekensis (Trägårdh, 1910) CU
WS England Denmark Austria Ukraine Crimea
Georgia
Tectocepheus velatus (Michael, 1880) RU
WS SS MS CY Belgium Denmark Georgia Kazakhstan
OTOCEPHEIDAE Balogh, 1961
Fissicepheus clavatus (Aoki, 1959) AM
OPPIOIDEA Grandjean, 1954
AUTOGNETIDAE Grandjean, 1960
Autogneta longilamellata (Michael,1885)SS
Autogneta willmanni (Dyrdowska, 1929) Belarus
Conchogneta tragardhi (Forsslund, 1947) WS
SS MS Belarus
Parautogneta golosovae Rjabinin, 1975 MS
Parautogneta silvatica Golosova, 1974 MS
CALEREMAEIDAE Grandjean, 1965
Caleremaeus monilipes (Michael, 1882) unknown
locality
Veloppia pulchra Hammer, 1952 USA(Alaska)
OPPIIDAE Grandjean, 1954
Berniniella bicarinata (Paoli, 1908) WS
Berniniella rafalski (Oplotna et Rajski, 1983) WS
MS
Cassioppia krivolutskyi Poltavskaja, 1994 WS
MS Kazakhstan
Lauroppia maritima (Willmann, 1929) WS
MS
Lauroppia neerlandica (Oudemans, 1900) WS
MS
Medioppia tuberculata (Bulanova-Zachvatkina, 1964) WS
Microppia minus (Paoli, 1908) WS
Microppia minutissima (Sellnick, 1950) WS
SS Kazakhstan
Moritzoppia keilbachi (Moritz, 1969) SS
MS
Moritzoppiella microdentata (Gordeeva et Grishina, 1991) WS
Tetroppia minuta (Gordeeva et Grishina, 1991) WS
SS
Tetroppia nikolskii (Gordeeva et Grishina, 1991) paratype!
WS
SS
Tetroppia nitens (Gordeeva et Grishina, 1991) WS
SS MS
Tetroppia praestans (Gordeeva et Grishina, 1991) WS
SS MS
Tetroppia similis (Gordeeva et Grishina, 1991) paratype!
WS
MS
Multioppia laniseta Moritz, 1966 SS Turkmenistan
Kazakhstan
Oppia denticulata (R. et G. Canestrini, 1882) paratype!
MS
Oppiella nova (Oudemans, 1902) WS SS MS
Azerbaijan
Kazakhstan
Oppiella trapesoida Grishina, 1981 paratype!
WS
SS
Ramusella fasciata (Paoli, 1908) Kazakhstan
SUCTOBELBIDAE Jacot, 1938
Allosuctobelba grandis (Paoli,1908) MS
Rhinosuctobelba dicerosa Woolley, 1969 SS
Suctobelbella acrana Moritz, 1970 WS SS
MS
Suctobelbella acutidens (Forsslund, 1941) WS
Suctobelbella forsslundi (Strenzke, 1950) WS
Suctobelbella hammeri (Krivolutsky, 1966) WS
SS MS
Suctobelbella nasalis (Forsslund, 1941) MS
Kazakhstan
Suctobelbella palustris (Forsslund, 1953) WS
MS
Suctobelbella prominens (Moritz, 1966) WS
SS
Suctobelbella singularis (Strenzke, 1950) WS
Suctobelbella subcornigera (Forsslund, 1941) WS
SS MS Kazakhstan
Suctobelbella subtrigona (Oudemans, 1900) WS
Suctobelbella tatarica (Krivolutsky, 1968) WS
Suctobelbella vera (Moritz, 1964) WS
THYRISOMIDAE Grandjean, 1954
Banksinoma lanceolata (Michael, 1885) WS
SS MS
Banksinoma setosa Rjabinin, 1974 SS
Kaszabobates kaszabi Balogh et Mahunka, 1965 RU
Kaszabobates olbiopolitanus Sergienko, 1980 paratype!
Ukraine
Pantalozetes paoli (Oudemans, 1913) WS
SS MS
HYDROZETOIDEA Grandjean, 1954
HYDROZETIDAE Grandjean, 1954
Hydrozetes confervae (Schrank,1781) WS
Hydrozetes parisiensis Grandjean, 1948 WS
Hydrozetes tamarae Tolstikov, 1996 paratype!
WS
LIMNOZETIDAE Grandjean, 1954
Limnozetes sphagni (Michael, 1884) WS
AMERONOTHROIDEA Willmann, 1931
AMERONOTHRIDAE Willmann, 1931
Ameronthrus dubinini Sitnikova, 1975 SS
CYMBAEREMAEOIDEA Sellnick, 1928
CYMBAEREMAEIDAE Sellnick, 1928
Scapheremaeus palustris (Sellnick, 1924) FS
Cuba
MICREREMIDAE Grandjean, 1954
Micreremus brevipes (Michael, 1888) WS
MS
Micreremus gracilior (Willmann, 1931) MS
Pterogasterina - Poronota
LICNEREMAEOIDEA Grandjean, 1931
LICNEREMAEIDAE Grandjean, 1931
Licneremaeus licnophorus (Michael, 1882) SS
MS
Licneremaeus prodigious Schuster, 1958 WS
SS Kazakhstan
PASSALOZETIDAE Grandjean,1954
Bipassalozetes elegans (Sitnikova, 1975) Kazakhstan
Bipassalozetes granulatus (Mihelcic, 1955) RU
Bipassalozetes intermedius (Mihelcic, 1954) SS
Bipassalozetes lineolatus (Sitnikova, 1975) Kazakhstan
Bipassalozetes perforatus (Berlese, 1910) SS
Bipassalozetes pilosus Bayartogtokh et Smelyansky, 2003 HOLOTYPE!
paratypes! SS
Bipassalozetes reticulates (Mihelcic, 1957) Kazakhstan
Bipassalozetes rugosus (Sitnikova, 1975) WS
Kazakhstan
Passalozetes africanus Grandjean, 1932 Kazakhstan
SCUTOVERTICIDAE Grandjean, 1954
Hypovertex borealis Rjabinin, 1984 KM
Hypovertex mirabilis Krivolutsky, 1969 WS
Latovertex laticuspis (Balogh et Mahunka, 1965) WS
SS
Scutovertes glandulosus Balogh et Mahunka, 1965 Ukraine
Scutovertex lanceolatus Hammer, 1952 Canada
Scutovertes minutus (C.L.Koch, 1836) Kazakhstan
Scutovertes ornatus Bulanova-Zachvatkina, 1964 RU
Scutovertes perforatus Sitnikova, 1975 SS
Scutovertes rugosus Mihelcic, 1975 SS
Kazakhstan
Scutovertes sculptus Michael, 1879 WS
SS Kazakhstan
Scutovertes serratus Sitnikova, 1975 SS
ORIPODOIDEA Jacot, 1925
HAPLOZETIDAE Grandjean, 1936
Haplozetes vindobonensis (Willmann, 1935) WS
SS MS Turkmenistan
Peloribates canadensis Hammer, 1952 Canada
Peloribates europaeus Willmann, 1935 WS
Peloribates pilosus Hammer, 1952 WS SS
Canada
Peloribates robustus Grishina, 1981 MS
ORIBATULIDAE Thor, 1929
Eporibatula prominens Bayartogtokh et Aoki, 1998 SS
Oribatula angustolamellata Iordanski, 1991 paratype!
Ukraine
Oribatula commutata Iordanski, 1991 paratype!
MS
Oribatula elegantissima Balogh et Mahunka, 1965 SS
Oribatula pallida Banks, 1906 WS SS MS
KU Kazakhstan
Oribatula robusta Iordanski, 1991 paratype!
RU
Oribatula sitnikavae Iordanski, 1991 paratype!
FS
Oribatula tibialis (Nicolet,1855) WS MS
Lithuania
Oribatula tibialis ssp. alata Iordanski, 1991 paratype!
RU
Oribatula variaporosa Iordanski, 1991 paratype!
RU
Paraleius leontonychus (Berlese, 1910)
MS
Simkinia elongata Krivolutsky, 1969 WS
Simkinia montana Krivolutsky et Grishina, 1970 SS
Simkinia turanica Krivolutsky, 1966 WS
Kazakhstan
Zygoribatula cognata (Oudemans, 1902) SS
Zygoribatula concina Iordanski, 1990 paratype!
Ukraine
Zygoribatula exilis (Nicolet, 1855) SS
Lithuania
Zygoribatula frisiae (Oudemans, 1900) RU
WS Kazakhstan
Zygoribatula propinquus (Oudemans, 1902) RU
SS MS
Zygoribatula terricola van der Hammen, 1952 RU
SS
PROTORIBATIDAE J. Balogh et P.Balogh, 1984
Liebstadia similis (Michael, 1888) WS
SS Georgia
Protoribates badensis Sellnick, 1928 unknown
locality
Protoribates capucinus (Berlese, 1908) WS
SS Turkmenistan
Protoribates divergens Mihelcic, 1955 SS
Kyrghyzstan
Protoribates variabilis Rajski, 1958 WS
SS
SCHELORIBATIDAE Grandjean, 1953
Scheloribates fimbriatus Thor, 1930 Turkmenistan
Scheloribates laevigatus (C.L.Koch, 1844) RU
Lithuania
Finland Belarus
Scheloribates latipes (C.L.Koch, 1844) Belarus
Scheloribates semidesertus Bulanova-Zachvatkina et Mahmudova,
1991 paratype! RU Turkmenistan
Uzbekistan
Topobates humeralis Subbotina, 1976 RU
WS
PARAKALUMMIDAE Grandjean, 1936
Neoribates aurantiacus (Oudemans, 1914) FS RU WS SS MS AM
KM Lithuania Georgia
Neoribates gracilis Trave, 1972 WS SS MS AM
Lithuania Belarus Ukraine
CERATOZETOIDEA Jacot, 1925
CERATOZETIDAE Jacot, 1925
Ceratozetes cisalpinus Berlese, 1908 RU
WS SS AM
Ceratozetes contiquus Jeleva, 1962 Crimea
Ceratozetes cuspidatus Jecot, 1939 Canada
Ceratozetes gracilis (Michael,1884) WS
Ceratozetes inupiag Behan-Pelletier, 1986 paratype!
Canada
Ceratozetes japonicus Aoki, 1960 SS
Ceratozetes kananaskis Mitchell, 1976 Canada
Ceratozetes kutchin Behan-Pelletier, 1986 paratype!
Canada
Ceratozetes laticuspidatus Menke, 1964 Turkmenistan
Ceratozetes macromediocris Shaldybina, 1970 RU
Ceratozetes mediocris Berlese, 1908 AM
Belarus
Ceratozetes minutissimus Willmann, 1951 RU
Kazakhstan
Ceratozetes oresbios Behan-Pelletier, 1984 Canada
Ceratozetes peritus Grandgean, 1951 SS
Ceratozetes sellnicki Rajski, 1958 WS
SS AM Kazakhstan
Ceratozetes spitsbergensis Thor, 1934 USA(Alaska)
Ceratozetes thienemanni Willmann, 1943 RU
WS MS
Ceratozetes virginicus (Banks, 1906) Canada
Cyrtozetes denaliensis Behan-Pelletier, 1985 Canada
Dentizetes rudentiger Hammer, 1952 Canada
Diapterobates arnoldii Krivolutsky, 1966 Kyrghyzstan
Diapterobates dubinini Shaldybina, 1971 SS
Diapterobates humeralis (Hermann, 1804) MS
Kazakhstan
Diapterobates notatus (Thorell, 1871) WS
SS MS
Diapterobates oblongus (C.L.Koch, 1879) WS
MS Kazakhstan
Diapterobates reticulatus (L.Koch, 1879) WS
SS MS
Diapterobates rotundocuspidatus Shaldybina, 1970 unknown
locality
Diapterobates rostralis Shaldybina, 1971 WS
Diapterobates siccatus Behan-Pelletier, 1986
paratype! USA(Alaska)
Diapterobates sitnikovae Shaldybina, 1970 SS
Diapterobates variabilis Hammer, 1955 WS
SS USA(Alaska)
Edwardzetes edwardsii (Nicolet, 1855) Ukraine
Fuscozetes fuscipes (C.L.Koch, 1844) unknown
locality
Fuscozetes pseudosetosus Shaldybina, 1979 SS
MS
Globozetes longipilus Sellnick, 1928 unknown
locality
Iugoribates gracilis Sellnick, 1944 USA(Alaska)
Laminizetes fortispinosus Behan-Pelletier, 1986 paratype!
Canada
Latilamellobates incisellus (Kramer, 1897) WS
SS MS
Latilamellobates naltschicki Shaldybina, 1971 RU
Melanozetes altaicus Shaldybina, 1969 SS
Melanozetes interruptus Willmann, 1953 SS
MS
Melanozetes longisetosus Hammer, 1952 USA(Alaska)
Melanozetes mollicomus (C.L.Koch, 1839) RU
Melanozetes orientalis Shaldybina, 1969 WS
SS MS
Melanozetes sellnicki (Hammer, 1952) WS
Melanozetes tanana Behan-Pelletier, 1986
paratype!
Canada
Neogymnobates luteus (Hammer, 1955) Canada
Sphaerozetes arcticus Hammer, 1952 MS
Canada
Sphaerozetes firthensis Behan-Pelletier, 1986
paratype! Canada
Sphaerozetes orbicularis (C.L.Koch, 1836) WS
MS
Sphaerozetes castaneus Hammer, 1955 Canada
Svalbardia paludicola Thor, 1930 WS MS
Canada
Trichoribates copperminensis Hammer, 1952 Canada
Trichoribates longipilis Willmannn, 1951 Ukraine
Trichoribates novus (Sellnick, 1928) WS
SS
Trichoribates ogilviensis Behan-Pelletier, 1986 Canada
Trichoribates polaris Hammer, 1953 Canada
Trichoribates punctatus Shaldybina, 1971 WS
Kazakhstan
Trichoribates striatus Hammer, 1952 Canada
Trichoribates trimaculatus (C.L.och, 1835) WS
SS
CHAMOBATIDAE Thor, 1838
Chamobates cuspidatus (Michael, 1884) WS
MS
Chamobates dentotutorii Shaldybina, 1969 Kazakhstan
Chamobates subglobosus (Oudemans, 1900) Ukraine
Xiphobates spinosus (Sellnick, 1928) unknown
locality
Xiphobates voigtsi (Oudemans, 1902) unknown
locality
EUZETIDAE Grandjean, 1954
Euzetes globulus (Nicolet, 1855) MS
HUMEROBATIDAE Grandjean, 1970
Humerobates translamellatus Shaldybina, 1971 WS
SS Kazakhstan
MYCOBATIDAE Grandjean, 1954
Minunthozetes pseudofusiger (Schweizer, 1922) RU
Mycobates bicornis (Strenzke, 1954) SS
MS
Mycobates monodactylus Schaldybina, 1970 SS
Mycobates patrius Schaldybina, 1970 WS
MS
Mycobates sarekensis (Trägårdh, 1910) SS
Pelopsis bifurcatus Ewing, (1909) Canada
Punctoribates grishinae Shaldybina, 1970 RU
WS
Punctoribates hexagonus Berlese, 1908 WS
Punctoribates manzanoensis Hammer, 1958 WS
Punctoribates meridianus Shaldybina, 1973 RU
Punctoribates minimus Shaldybina, 1969 RU
WS Azerbaijan Kazakhstan
Punctoribates punctum (C.L.Koch, 1839) RU
Ukraine
Cuba
Punctoribates sellnicki (Willmann, 1928) WS
Punctoribates sphaericus Shaldybina, 1987 paratype!
WS
Punctoribates sibiricus Grishina, 1984 paratype!
WS
Punctoribates zachvatkini Shaldybina, 1969 WS
Zachvatkinibates conjunctus Shaldybina, 1973 SS
ZETOMIMIDAE Shaaldybina, 1966
Heterozetes palustris Willmann, 1917 WS
Zetomimus furcata (Pearce et Warburton, 1905) WS
PHENOPELOPOIDEA Petrunkevitch, 1955
PHENOPELOPIDAE Petrunkevitch, 1955
Eupelops acromios (Hermann, 1804) WS SS
MS
Eupelops nepotulus (Berlese, 1916) WS
SS MS Kazakhstan
Eupelops occultus (C.L.Koch, 1836) WS
Eupelops plicatus (C.L.Koch, 1835) WS
SS
Eupelops subuliger (Berlese, 1916) WS
MS
Eupelops torulosus (C.L.Koch, 1836) WS
MS
Peloptulus gibbus Mihelcic, 1957 WS SS
Kazakhstan
Peloptulus phaenotus (C.L.Koch, 1844) WS
SS
Peloptulus reticulatus Mihelcic, 1957 WS
SS Kazkhstan
Propelops canadensis (Hammer, 1952) WS
SS Kazakhstan Canada
Propelops groenlandicus (Sellnick, 1944) AM
Canada
ORIBATELLOIDEA Jacot, 1925
ORIBATELLIDAE Jacot, 1925
Kunstella foveolata Krivolutsky, 1974 SS
Oribatella byzovae Krivolutsky, 1974 SS
Oribatella calcarata (C.L.Koch, 1835) WS
SS
Oribatella quadricornuta (Michael, 1880) WS
SS
Oribatella reticulata Berlese, 1916 WS
Kazakhstan
Oribatella shaldybinae Rjabinin, 1974 SS
ACHIPTERIOIDEA Thor, 1929
ACHIPTERIIDAE Thor, 1929
Achipteria coleoptrata (Linnaeus, 1758) FS
Kazakhstan
Achipteria nitens (Nicolet, 1855) WS SS
Anachipteria latitecta (Berlese, 1908) WS
Kazakhstan
Parachipteria bella (Sellnick, 1928) SS
Parachipteria punctata (Nicolet, 1855) Ukraine
Parachipteria sibirica Krivolutsky et Grishina, 1970 paratype!
MS
Parachipteria willmanni van der Hammen, 1952 WS
SS MS
Tectoribates ornatus (Schuster, 1958) WS
SS Kazakhstan
TEGORIBATIDAE Grandjean, 1954
Lepidozetes chernovi Rjabinin, 1974 WS
SS Kazakhstan
Lepidozetes latipilosus Hammer, 1952 SS
Canada
Lepidozetes singularis Berlese, 1910 WS
SS MS
Scutozetes lanceolatus Hammer, 1952 SS
MS
Tegoribates latirostris (C.L. Koch, 1844) WS
SS
Umbellozetes fuscus Krivolutsky, 1969 SS
GALUMNOIDEA Jacot, 1925
GALUMNIDAE Jacot, 1925
Acrogalumna longipluma (Berlese,1904) CU
AM Austria Bulgaria Ukraine Georgia
Lithuania Cuba
Angulogalumna asiatica Grishina, 1981
paratipe! SS AM
Galumna alata (Hermann, 1804)
Kirghizia Kazakhstan
Galumna banksi Jacot, 1929 USA
Galumna cribriger (Berlese, 1916) USA
Galumna dimorpha Krivolutskaja, 1952 RU
SS MS
Galumna flagellata Willmann, 1925 WS AM
Kazakhstan
Galumna hawaiensis Jacot, USA
Galumna hudsoni Hammer, 1952 Canada
Galumna lanceata Oudemans, 1900 FS RU
CU WS SS MS AM Ukraine Crimea
Galumna minuta (Ewing, 1909)
Galumna obvia (Berlese,1914) FS RU CU
AM Yugoslavia Bulgaria Belarus
Galumna rossica Sellnick, 1926 WS SS MS
AM Crimea
Galumna tarsipennata Oudemans, 1914 RU
AM Belarus Ukraine
Galumna virginiensis Jacot, 1929 USA
Galumna zachvatkini Grishina,1981 RU Bulgaria
Ukraine
Holokalumma coloradensis Jacot, 1929 USA
Notogalumna nortoni Ramani et Haq India
Orthogalumna terebrantis Wallwork, 1965 USA
Pergalumna altera (Oudemans, 1915) Ukraine
Kazakhstan
Pergalumna curva (Ewing, 1907) Ewing, 1907 USA
Pergalumna duplicata ssp. nipponica Aoki, 1966 AM
Pergalumna emarginata (Banks, 1895) USA
Pergalumna flagellifera (Jacot, 1929) USA
Pergalumna formicaria (Berlese, 1914) Canada
Pergalumna minor (Willmann, 1928) CU Bulgaria
Pergalumna myrmophila (Berlese, 1914) CU
Ukraine
Crimea Georgia
Pergalumna nervosa (Berlese, 1914) FS
RU CU WS MS SH Denmark Austria Bulgaria Finland
Ukraine Crimea Georgia
Pergalumna willmanni (Zachvatkin, 1953) FS
RU WS SS MS AM
Pilogalumna allifera (Oudemans, 1915) RU
CU WS Bulgaria Ukraine Crimea Georgia
Pilogalumna allifera montana Grishina et Sergienko, 1978 paratype!
Crimea
Pilogalumna cozadensis Nevin, 1976 Canada
Pilogalumna tenuiclava (Berlese, 1908) CU
WS SS MS Lithuania Ukraine Georgia Mongolia
Psammogalumna hungarica (Sellnick, 1925) WS
Crimea
Psammogalumna thysanura (Krivolutskaja, 1952) WS
Crimea
Kazakhstan