Carabinae, Bembidiini
collection of Siberian Zoological Museum
(curator - Roman Yu. Dudko)

materials from different regions of Russia are encoded as: FS RU CU UR WS SS MS CY NE NF KM AM SH KU

Asaphidion austriacum Schweiger, 1975 CU
Asaphidion caraboides (Schrank, 1781)
Asaphidion caraboides caraboides (Schrank, 1781) Ukraine
Asaphidion caraboides varipes (Motschulsky, 1856) CU
Asaphidion cupreum Andrewes, 1925 SS
Asaphidion curtum Heyden, 1870 France
Asaphidion flavicorne (Solsky, 1874) Kazakhstan
Asaphidion flavipes (Linnaeus, 1761) RU WS
Asaphidion pallipes (Duftschmid, 1812) RU CU WS SS MS
Asaphidion semilucidum (Motschulsky, 1862) AM
Asaphidion stierlini Heyden, 1880 France
Asaphidion transcaspicum Semenov, 1889 Kazakhstan
Asaphidion ussuriense Jedlichka, 1965 AM
Asaphidion indicum Chaudoir India
Ocys (Ocys) harpaloides (Serville, 1821) France
Ocys (Ocys) quinquestriatus (Gyllenhal, 1810) France
Bembidion (Microserrullula) quadricolle (Motschulsky, 1844) CU Turkmenistan
Bembidion (Bracteon) argenteolum (Ahrens, 1812) RU WS SS
Bembidion (Bracteon) foveum Motschulsky, 1844 WS SS
Bembidion (Bracteon) lapponicum Zetterstedt, 1828 WS SS AM NF
Bembidion (Bracteon) litorale (Olivier, 1790) RU WS
Bembidion (Bracteon) conicolle Motschulsky, 1844 SS AM
Bembidion (Bracteon) stenoderum H.Bates, 1873 AM
Bembidion (Bracteon) zephyrum Fall, 1910 USA
Bembidion (Bracteon) inaequale Say, 1823 USA
Bembidion (Bracteon) lorquini Chaudoir, 1868 USA
Bembidion (Ochthedromus) americanum Dejean, 1831 USA
Bembidion (Odontium) striatum (Fabricius, 1792) WS SS Mongolia
Bembidion (Odontium) chloropus H.Bates, 1883 KU
Bembidion (Odontium) confusum Hayward, 1897 USA
Bembidion (Eudromus) obliquulum Leconte, 1859 USA
Bembidion (Euryrtachelus) laevibase (Reitter, 1902) Kazakhstan
Bembidion (Euryrtachelus) vitiosum Gemminger et Harold, 1868 WS SS
Bembidion (Neja) submutatum Netolitzky, 1911 Tadzhikistan
Bembidion (Chlorodium) almum J.Sahlberg, 1900
Bembidion (Chlorodium) almum almum J.Sahlberg, 1900 RU Kazakhstan
Bembidion (Chlorodium) almum protalmum Netolitzky, 1933 SS
Bembidion (Chlorodium) difforme (Motschulsky, 1844) SS
Bembidion (Chlorodium) luridicorne Solsky, 1874 Turkmenistan
Bembidion (Chlorodium) luteipes (Motschulsky, 1844) Uzbekistan
Bembidion (Chlorodium) pygmaeum (Fabricius, 1792) Germany
Bembidion (Metallina) elevatum (Motschulsky, 1844)
Bembidion (Metallina) elevatum lamprosimile Netolitzky, 1942 AM
Bembidion (Metallina) lampros (Herbst, 1784) RU WS SS Ukraine Georgia Kazakhstan
Bembidion (Metallina) properans (Stephens, 1829) RU WS SS Kazakhstan
Bembidion (Phyla) obtusum Serville, 1821 Germany
Bembidion (Phyla) incommodum Netolitzky, 1926 CU
Bembidion (Phyla) tethys Netolitzky, 1926 Spain
Bembidion (Phyla) demidenkoae Dudko, 1999 HOLOTYPE! SS
Bembidion (Princidium) punctulatum Drapiez, 1820 Latvia
Bembidion (Paraprincidium) ruficolle (Panzer, 1797) WS
Bembidion (Testedium) bipunctatum (Linnaeus, 1761)
Bembidion (Testedium) bipunctatum bipunctatum (Linnaeus, 1761) SS Kazakhstan
Bembidion (Testedium) bipunctatum pyritosum Rossi Italy
Bembidion (Testedium) bipunctatum gracile Abejar Spain
Bembidion (Testedium) bipunctatum rugiceps Chaudoir, 1846 Turkey
Bembidion (Testedium) bipunctatum tectimundi Netolitzky, 1943 Kyrghyzstan
Bembidion (Testedium) laetum Brulle, 1839 Morocco
Bembidion (Actedium) pallidipenne (Illiger, 1801) France
Bembidion (Notaphus) obliquum Sturm, 1825 WS SS AM
Bembidion (Notaphus) pedestre (Motschulsky, 1844) WS SS
Bembidion (Notaphus) semipunctatum (Donovan, 1806) RU WS SS AM SH
Bembidion (Notaphus) varium (Olivier, 1795) RU WS SS AM SH Latvia
Bembidion (Notaphus) graphicum Casey, 1918 USA
Bembidion (Notaphus) approximatum Leconte, 1852 USA
Bembidion (Eupetedromus) sibiricum Dejean, 1831 SS
Bembidion (Eupetedromus) incrematum Leconte, 1860 Canada
Bembidion (Notaphemphanes) ephippium (Marsham, 1802) RU
Bembidion (Notaphocampa) niloticum Dejean, 1831
Bembidion (Notaphocampa) niloticum hamatum Kolenati, 1845 Turkmenistan Tadzhikistan
Bembidion (Philochtus) aeneum Germar, 1824 Netherlands
Bembidion (Philochtus) biguttatum (Fabricius, 1779) RU WS Kazakhstan
Bembidion (Philochtus) ellipticocurtum Netolitzky, 1935 Armenia
Bembidion (Philochtus) inoptatum Schaum, 1857 Austria Ukraine
Bembidion (Philochtus) iricolor Bedel, 1885 France
Bembidion (Philochtus) guttula (Fabricius, 1792) RU WS SS
Bembidion (Philochtus) lunulatum (Fourcroy, 1785) France
Bembidion (Philochtus) mannerheimi C.R.Sahlberg, 1834 WS SS Kazakhstan
Bembidion (Philochtus) paganettii Netolitzky, 1914 Spain
Bembidion (Philochtus) pallidiveste Carrett, 1906 RU
Bembidion (Lionepha) erasum Leconte, 1859 USA
Bembidion (Emphanes) azurescens Dalla Torre, 1877 France
Bembidion (Emphanes) latiplaga Chaudoir, 1850 RU WS
Bembidion (Emphanes) gobiense Jedlichka, 1964 SS
Bembidion (Emphanes) minimum (Fabricius, 1792) RU WS SS
Bembidion (Emphanes) normannum Dejean, 1831 France
Bembidion (Emphanes) rivulare Dejean, 1831
Bembidion (Emphanes) rivulare rivulare Dejean, 1831 France
Bembidion (Emphanes) rivulare euxinum Apfelbeck, 1904 RU SS
Bembidion (Emphanes) axillare (Motschulsky, 1844) WS SS
Bembidion (Emphanes) quadriplagiatum (Motschulsky, 1844) WS SS
Bembidion (Emphanes) tenellum Erichson, 1837
Bembidion (Emphanes) tenellum tenellum Erichson, 1837 RU WS France
Bembidion (Talanes) aspericolle (Germar, 1812) WS
Bembidion (Talanes) subfasciatum Chaudoir, 1850 RU
Bembidion (Leja) articulatum (Panzer, 1796) RU WS SS AM Hungary Kazakhstan
Bembidion (Leja) octomaculatum (Goeze, 1777) RU WS AM Kazakhstan
Bembidion (Leja) maculatum Dejean, 1831 Croatia
Bembidion (Trepanedoris) doris (Panzer, 1797) RU WS SS
Bembidion (Trepanedoris) fortestriatum Motschulsky, 1845 Canada
Bembidion (Semicampa) amoenum R.F.Sahlberg, 1844 AM
Bembidion (Semicampa) chaudoiri Chaudoir, 1850 WS
Bembidion (Semicampa) dormeyeri Reitter, 1897 SS
Bembidion (Semicampa) gassneri Netolitzky, 1922 WS
Bembidion (Semicampa) gilvipes Sturm, 1825 RU WS Kazakhstan
Bembidion (Semicampa) guttulatum Chaudoir, 1850 CU
Bembidion (Semicampa) mandarin Netolitzky, 1939 SS
Bembidion (Semicampa) schuppelii Dejean, 1831 RU WS SS Kazakhstan
Bembidion (Semicampa) variola Netolitzky, 1810 Tadzhikistan
Bembidion (Diplocampa) assimile Gyllenhal, 1810 CU WS Kazakhstan
Bembidion (Diplocampa) bisulcatum Chaudoir, 1844 WS
Bembidion (Diplocampa) fumigatum (Duftschmid, 1812) SS
Bembidion (Diplocampa) transparens (Gebler, 1829)
Bembidion (Diplocampa) transparens transparens (Gebler, 1829) WS SS Kazakhstan
Bembidion (Diplocampa) transparens prostratum (Motschulsky, 1844) SS AM China
Bembidion (Diplocampa) blandulum Netolitzky, 1910 SS
Bembidion (Bembidion) humerale Sturm, 1825 WS
Bembidion (Bembidion) quadrimaculatum (Linnaeus, 1761)
Bembidion (Bembidion) quadrimaculatum quadrimaculatum (Linnaeus, 1761) RU WS MS SS Kazakhstan
Bembidion (Bembidion) quadrimaculatum mandli Netolitzky, 1932 SS AM
Bembidion (Bembidion) quadripustulatum (Serville, 1821) SS Tadzhikistan
Bembidion (Nepha) genei Kuster, 1847
Bembidion (Nepha) genei genei Kuster, 1847 France
Bembidion (Nepha) genei illigeri Netolitzky, 1914 RU France
Bembidion (Nepha) menetriesi Kolenati, 1845
Bembidion (Nepha) menetriesi menetriesi Kolenati, 1845 Turkmenistan
Bembidion (Nepha) menetriesi retipenne J.Mueller, 1918 Turkey
Bembidion (Nepha) rufimacula J.Mueller, 1918 Turkey
Bembidion (Nepha) seriatum (Motschulsky, 1844) CU
Bembidion (Nepha) tetrasemum Chaudoir, 1846 Georgia
Bembidion (Nepha) tetragrammum Chaudoir, 1846 CU
Bembidion (Nepha) alluaudi Antoine, 1939 Morocco
Bembidion (Nepha) fortunatum Wollaston, 1871 Spain(Canarias)
Bembidion (Nepha) callosum Kuster, 1847
Bembidion (Nepha) callosum subconnexum Monte France
Bembidion (Nepha) subcallosum Wollaston, 1864 Spain(Canarias)
Bembidion (Bembidionetolitzkya) atrocoeruleum (Stephens, 1829) France
Bembidion (Bembidionetolitzkya) coeruleum Serville, 1821
Bembidion (Bembidionetolitzkya) coeruleum coeruleum Serville, 1821 France
Bembidion (Bembidionetolitzkya) coeruleum scelio Antoine Morocco
Bembidion (Bembidionetolitzkya) varicolor (Fabricius, 1803)
Bembidion (Bembidionetolitzkya) varicolor agurense Lutshnik, 1937 CU
Bembidion (Bembidionetolitzkya) conforme Dejean, 1831 France
Bembidion (Bembidionetolitzkya) ascendens K.Daniel, 1902 Austria
Bembidion (Bembidionetolitzkya) fasciolatus (Duftschmid, 1812) France
Bembidion (Bembidionetolitzkya) bactrianum K.Daniel, 1902 Tadzhikistan
Bembidion (Bembidionetolitzkya) astrabadense (Mannerheim, 1844)
Bembidion (Bembidionetolitzkya) astrabadense transcaucasicum Lutshnik, 1939 Turkey
Bembidion (Bembidionetolitzkya) tibiale (Duftschmid, 1812) Italy Germany Ukraine
Bembidion (Bembidionetolitzkya) cyaneum Chaudoir, 1846 CU
Bembidion (Bembidionetolitzkya) depressum Ménétriès, 1832 CU
Bembidion (Bembidionetolitzkya) complanatum Heer, 1837 France
Bembidion (Bembidionetolitzkya) relictum Apfelbeck, 1904 CU
Bembidion (Bembidionetolitzkya) geniculatum Heer, 1837
Bembidion (Bembidionetolitzkya) geniculatum geniculatum Heer, 1837 France Ukraine
Bembidion (Bembidionetolitzkya) geniculatum nevadense Jeannel Spain
Bembidion (Bembidionetolitzkya) geniculatum dilutipes Deville, 1806 France
Bembidion (Bembidionetolitzkya) kartalinicum Lutshnik, 1937 CU
Bembidion (Bembidionetolitzkya) piceocyaneum Solsky, 1874 Kyrghyzstan
Bembidion (Plataphus) altaicum (Gebler, 1833) SS Kazakhstan
Bembidion (Plataphus) planatum Leconte, 1848 SS
Bembidion (Plataphus) coelestinum (Motschulsky, 1844) SS
Bembidion (Plataphus) flebile Casey, 1918 USA
Bembidion (Plataphus) gebleri (Gebler, 1833)
Bembidion (Plataphus) gebleri gebleri (Gebler, 1833) SS Kazakhstan
Bembidion (Plataphus) gebleri persuasum Netolitzky, 1938 SS AM
Bembidion (Plataphus) gebleri turbatum Casey, 1908 USA
Bembidion (Plataphus) gratiosum Casey, 1918 USA
Bembidion (Plataphus) infuscatipenne Netilitzky, 1938 AM
Bembidion (Plataphus) hyperboraeorum Munster, 1923 WS
Bembidion (Plataphus) planiusculum Mannerheim, 1843 USA
Bembidion (Plataphus) planum R.F.Sahlberg, 1844 USA
Bembidion (Plataphus) prasinum (Duftschmid, 1812) WS Finland
Bembidion (Plataphus) stillaguamish Hatsch, 1950 USA
Bembidion (Plataphus) asiaticum Jedlichka, 1965 SS AM Kazakhstan
Bembidion (Plataphodes) aeruginosum (Gebler, 1833) SS Kazakhstan
Bembidion (Plataphodes) arcticum Lindroth, 1963 SS Kazakhstan
Bembidion (Plataphodes) complanulum Mannerheim, 1853 SS
Bembidion (Plataphodes) difficile (Motschulsky, 1844) UR WS SS Kazakhstan
Bembidion (Plataphodes) farrarae Hatsch, 1950 USA
Bembidion (Plataphodes) fellmanni (Mannerheim, 1823) WS SS Kazakhstan
Bembidion (Plataphodes) haruspex Casey, 1918 USA
Bembidion (Plataphodes) incertum Motschulsky, 1845 USA
Bembidion (Plataphodes) tetraporum H.Bates, 1883 AM SH
Bembidion (Hirmoplataphus) hirmocoelum Chaudoir, 1850 WS SS AM SH Kazakhstan
Bembidion (Hirmoplataphus) concolor Kirby, 1837 USA
Bembidion (Hirmoplataphus) recticolle Leconte, 1863 USA
Bembidion (Hirmoplataphus) quadrulum Leconte, 1861 USA
Bembidion (Trichoplataphus) hasti C.R.Sahlberg, 1827 WS SS
Bembidion (Trichoplataphus) virens Gyllenhal, 1827 WS Finland
Bembidion (Trichoplataphus) deplanatum A.Morawitz, 1862 SS
Bembidion (Trichoplataphus) lissonotum H.Bates, 1873 AM
Bembidion (Lymnaeoperyphus) quadriimpressum Motschulsky, 1860 SH KU
Bembidion (Euperyphus) fulvipes Sturm, 1827 Austria
Bembidion (Euperyphus) combustum Ménétriès, 1832
Bembidion (Euperyphus) combustum combustum Ménétriès, 1832 Turkey
Bembidion (Ocydromus) obscurellum (Motschulsky, 1844)
Bembidion (Ocydromus) obscurellum turanicum Csiki, 1928 WS SS Kazakhstan China
Bembidion (Ocydromus) insidiosum Solsky, 1874
Bembidion (Ocydromus) insidiosum insidiosum Solsky, 1874 SS Kazakhstan
Bembidion (Ocydromus) andreae (Fabricius, 1787)
Bembidion (Ocydromus) andreae atlantis Antoine Morocco
Bembidion (Ocydromus) andreae bualei Jacquelin du Val, 1852 France Ukraine
Bembidion (Ocydromus) andreae polonicum J. Mueller, 1830 WS SS
Bembidion (Ocydromus) femoratum Sturm, 1825
Bembidion (Ocydromus) femoratum femoratum Sturm, 1825 WS SS Kazakhstan
Bembidion (Ocydromus) fluviatile Dejean, 1831 WS
Bembidion (Ocydromus) testaceum Duftschmid, 1812 France Ukraine
Bembidion (Ocydromus) petrosum Gebler, 1833 WS SS Kazakhstan
Bembidion (Ocydromus) morawitzi Csiki, 1928 KU SH
Bembidion (Ocydromus) subcostatum (Motschulsky, 1850) Turkey
Bembidion (Ocydromus) tetracolum Say, 1823 WS France
Bembidion (Ocydromus) bruxellense Wesmael, 1835 FS UR WS SS Kazakhstan
Bembidion (Ocydromus) captivorum Netolitzky, 1943 SS AM
Bembidion (Ocydromus) poppii Netolitzky, 1914 SS
Bembidion (Ocydromus) rupicola Kirby, 1837 USA
Bembidion (Ocydromus) ripicola Dufour, 1820 France
Bembidion (Ocydromus) hispanicum Dejean, 1831 Morocco
Bembidion (Ocydromus) straussi Netolitzky, 1910
Bembidion (Ocydromus) straussi iridipiceum Fassati, 1957 Kazakhstan
Bembidion (Ocydromus) altestriatum Netolitzky, 1934 SS Kazakhstan
Bembidion (Ocydromus) amnicola J.Sahlberg, 1900 Turkmenistan
Bembidion (Ocydromus) sajanum Shilenkov, 1995 SS
Bembidion (Ocydromus) infuscatum Dejean, 1831 SS Mongolia
Bembidion (Ocydromus) pseudoinfuscatum Shilenkov, in litt. paratype! SS
Bembidion (Ocydromus) ovale Motschulsky, 1844 SS
Bembidion (Ocydromus) kazakhstanicum Kryzhanovskij, 1979 Kazakhstan
Bembidion (Ocydromus) obenbergeri Lutshnik, 1928 SS Kazakhstan
Bembidion (Ocydromus) semilunium Netolitzky, 1914 SS AM China
Bembidion (Ocydromus) smirnovi Kryzhanovskij, 1979 SS
Bembidion (Ocydromus) umiatense Lindroth, 1963 NF
Bembidion (Ocydromus) ustum Quensel, 1806 Tadzhikistan
Bembidion (Ocydromus) pulcherrimum Motschulsky, 1850 CU
Bembidion (Ocydromus) mckinleyi Fall, 1926
Bembidion (Ocydromus) mckinleyi scandicum Lindroth, 1943 WS SS Kazakhstan
Bembidion (Ocydromus) ovalipenne Solsky, 1874 Kazakhstan
Bembidion (Ocydromus) consummatum H.Bates, 1873 SS
Bembidion (Ocydromus) transversale Dejean, 1831
Bembidion (Ocydromus) transversale perspicuum Leconte, 1848 Canada
Bembidion (Ocydromus) siculum Dejean, 1831
Bembidion (Ocydromus) siculum certans Netolitzky Morocco
Bembidion (Ocydromus) atlanticum Wollaston, 1854
Bembidion (Ocydromus) atlanticum megaspilum F. Walker, 1871 Morocco
Bembidion (Ocydromus) decorum (Zenker in Panzer, 1801)
Bembidion (Ocydromus) decorum decorum (Zenker in Panzer, 1801) France
Bembidion (Ocydromus) decorum caraffai Deville, 1807 France
Bembidion (Ocydromus) decorum subconvexum K.Daniel, 1902 CU
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850) SH
Bembidion (Ocydromus) modestum (Fabricius, 1801) Ukraine
Bembidion (Ocydromus) saxatile Gyllenhal, 1827
Bembidion (Ocydromus) saxatile flavipalpe Netolitzky, 1930 Kyrghyzstan
Bembidion (Ocydromus) saxatile fuscomaculatum (Motschulsky, 1844) WS SS Kazakhstan
Bembidion (Ocydromus) scopulinum (Kirby, 1837) WS SS AM
Bembidion (Ocydromus) zolotarewi Reitter, 1910 CU
Bembidion (Ocydromus) praeustum Dejean, 1831
Bembidion (Ocydromus) praeustum praeustum Dejean, 1831 RU Yugoslavia Greece
Bembidion (Ocydromus) praeustum fauveli Ganglbauer, 1892 France
Bembidion (Ocydromus) kiritschenkoi Michailov, 1984 Kazakhstan
Bembidion (Ocydromus) marginipenne Solsky, 1874 Kazakhstan
Bembidion (Ocydromus) parsorum Netolitzky, 1934 Armenia
Bembidion (Ocydromus) dalmatinum Dejean, 1831
Bembidion (Ocydromus) dalmatinum dalmatinum Dejean, 1831 RU Greece
Bembidion (Ocydromus) dalmatinum haupti Reitter, 1908 Armenia
Bembidion (Ocydromus) dalmatinum lacrimans Netolitzky, 1935 Turkey
Bembidion (Ocydromus) latinum Netolitzky, 1911 France
Bembidion (Ocydromus) weiratherianum Netolitzky Turkey
Bembidion (Ocydromus) dudichi Csiki, 1928 Morocco
Bembidion (Ocydromus) fraxator Ménétriès, 1832 Georgia
Bembidion (Ocydromus) milleri Duval, 1851 Germany
Bembidion (Ocydromus) deletum Serville, 1821 SS France Kazakhstan
Bembidion (Ocydromus) brunnicorne Dejean, 1831 CU
Bembidion (Ocydromus) olegleonidovici Fassati, 1990 CU
Bembidion (Ocydromus) grapei Gyllenhal, 1827 WS AM Kazakhstan
Bembidion (Ocydromus) yuconum Fall, 1929 SS Kazakhstan
Bembidion (Ocydromus) platynoides Hayward, 1897 USA
Bembidion (Ocydromus) alticola Fiori, 1903 Italy
Bembidion (Ocydromus) lacunarium Zimmermann, 1869 USA
Bembidion (Ocydromus) adygorum Belousov et Sokolov, 1996 paratypes! Georgia
Bembidion (Ocydromus) amurense (Motschulsky, 1859) SS AM
Bembidion (Ocydromus) dauricum (Motschulsky, 1844) SS CY Kazakhstan
Bembidion (Liocosmius) mundum Leconte, 1852 USA
Bembidion (Liocosmius) horni Hayward, 1897 USA
Bembidion (Testediolum) corsicum Csiki France
Bembidion (Testediolum) jackueti Jeannel France
Bembidion (Testediolum) montanum Rambur Spain
Bembidion (Testediolum) kokandicum Solsky, 1874 SS
Bembidion (Testediolum) validum Netolitzky, 1920 Uzbekistan
Bembidion (Testediolum) pyrenaeum Dejean, 1831 France
Bembidion (Testediolum) commotum Casey, 1918 USA
Bembidion (Pamirium) platypterum Solsky, 1874 Turkmenistan
Bembidion (Synechostictus) ruficorne Sturm 1825 Spain
Bembidion (Synechostictus) elongatum Dejean, 1831 France
Bembidion (Synechostictus) nordmanni Chaudoir, 1844 CU
Bembidion (Synechostictus) multisulcatum Reitter, 1890 CU
Bembidion (Synechostictus) cribrum Duval, 1851 France
Bembidion (Synechostictus) atroviolaceus Dufour, 1820
Bembidion (Synechostictus) atroviolaceus atroviolaceus Dufour, 1820 France Ukraine
Bembidion (Pseudolimnaeum) lederi Reitter, 1888 CU
Bembidion (Pseudolimnaeum) inustum Duval, 1857 Germany
Bembidion adductum Casey, 1918 USA
Bembidion obtusangulum Leconte, 1863 USA
Bembidion scudderi Leconte, 1878 USA


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