Carabinae, Harpalini
collection of Siberian Zoological Museum
(curator - Roman Yu. Dudko)

materials from different regions of Russia are encoded as: FS RU CU UR WS SS MS CY NE NF KM AM SH KU

Anisodactylus (Anisodactylus) binotatus (Fabricius, 1787) RU WS SS Kazakhstan
Anisodactylus (Anisodactylus) nemorivagus (Duftschmid, 1812) RU SS Ukraine Georgia
Anisodactylus (Anisodactylus) signatus (Panzer, 1797) RU CU WS SS AM SH KU Kazakhstan
Anisodactylus (Hexatrichus) poeciloides (Stephens, 1828)
Anisodactylus (Hexatrichus) poeciloides pseudaeneus Dejean, 1828 RU WS SS Azerbaijan Uzbekistan Kazakhstan
Diachromus germanus (Linnaeus, 1758) Georgia Azerbaijan
Bradycellus (Bradycellus) caucasicus (Chaudoir, 1846) RU WS SS Kazakhstan
Bradycellus (Tachycellus) laevicollis Poppius, 1907 AM
Bradycellus (Tachycellus) glabratus (Reitter, 1894) SS AM
Bradycellus (Tachycellus) subditus (Lewis, 1879) AM
Bradycellus (Tachycellus) glabratulus Lafer, 1989 AM
Bradycellus (Tachycellus) curtulus (Motschulsky, 1860) KU
Dicheirotrichus (Dicheirotrichus) abdominalis (Motschulsky, 1844) SS
Dicheirotrichus (Dicheirotrichus) ustulatus (Dejean, 1829) WS Kazakhstan
Dicheirotrichus (Dicheirotrichus) desertus (Motschulsky, 1850) WS SS Kazakhstan
Dicheirotrichus (Cardiostenus) medvedevi (Kabak et Kataev, 1993) Kyrghyzstan
Dicheirotrichus (Cardiostenus) microderus Solsky, 1874 Turkmenistan
Dicheirotrichus (Oreoxenus) mannerheimi (R.F.Sahlberg, 1844)
Dicheirotrichus (Oreoxenus) mannerheimi ponojensis (J.Sahlberg, 1875) SS Kazakhstan
Dicheirotrichus (Trichocellus) angularis (Reitter in Tschitscherine, 1899) SS
Dicheirotrichus (Trichocellus) punctatellus (Reitter, 1894) SH
Dicheirotrichus (Trichocellus) rufithorax (C.R.Sahlberg, 1827) WS Kazakhstan
Dicheirotrichus (Trichocellus) cognatus (Gyllenhal, 1827) RU WS SS
Dicheirotrichus (Trichocellus) externepunctatus Reitter in Tschitscherine, 1899 Kazakhstan
Dicheirotrichus (Trichocellus) placidus (Gyllenhal, 1827) WS
Lioholus jedlickai Lafer, 1989 AM
Stenolophus (Stenolophus) teutonus (Schrank, 1781) Italia Moldova
Stenolophus (Stenolophus) persicus Mannerheim, 1844 CU Georgia Turkmenistan Uzbekistan Tadzhikistan
Stenolophus (Stenolophus) discophorus (Fischer von Waldheim, 1823) RU WS Moldova Kazakhstan
Stenolophus (Stenolophus) skrimshiranus Stephens, 1828 CU Ukraine
Stenolophus (Stenolophus) propinquus A.Morawitz, 1862 AM
Stenolophus (Stenolophus) mixtus (Herbst, 1784) RU CU WS SS Hungary Ukraine Kazakhstan
Stenolophus (Stenolophus) proximus Dejean, 1829 RU Ukraine
Stenolophus (Stenolophus) castaneipennis H.Bates, 1873 AM
Stenolophus (Stenolophus) ussuricus Kataev et Dudko, 1997 HOLOTYPE! paratypes! AM
Stenolophus (Egadroma) marginatus (Dejean, 1829) Turkmenistan Tadzhikistan
Stenolophus (Egadroma) baiaurae And. Tadzhikistan
Stenolophus (Egadroma) quinquepustulata Westwood Thailand
Hemiaulax morio (Ménétriès, 1832) Tadzhikistan Kazakhstan
Loxoncus cyanescens (Hope, 1845) AM
Loxoncus procerus (Schaum, 1858) Azerbaijan
Acupalpus (Ancylostria) interstitialis Reitter, 1884 Moldova Georgia
Acupalpus (Acupalpus) flavicollis (Sturm, 1825) UR
Acupalpus (Acupalpus) meridianus (Linnaeus, 1767) RU WS SS Ukraine Kazakhstan
Acupalpus (Acupalpus) suturalis Dejean, 1829 Moldova
Acupalpus (Acupalpus) elegans (Dejean, 1829) RU WS Kazakhstan
Acupalpus (Acupalpus) parvulus (Sturm, 1825) RU WS SS Tadzhikistan Kazakhstan
Acupalpus (Acupalpus) maculatus Schaum, 1860 Hungary
Acupalpus (Acupalpus) flaviceps Motschulsky, 1850 RU Turkmenia Tadzhikistan
Acupalpus (Acupalpus) luteatus (Duftschmid, 1812) Moldova
Acupalpus (Acupalpus) exiguus (Dejean, 1829) RU WS SS Moldova Kazakhstan
Acupalpus (Setacupalpus) hilaris Tschitscherine, 1899 AM
Acupalpus (Palcuapus) inornatus H.Bates, 1873 AM
Anthracus consputus (Duftschmid, 1812) RU WS SS
Anthracus longicornis (Schaum, 1857) Moldova
Platymetopus flavilabris (Fabricius, 1798) Thailand
Dioryche indochinensis Bates Vietnam
Dioryche torta Mac Leag Vietnam
Chydaeus planicollis Andrewes Nepal
Pseudognathophanus punctilabris Mac Leag Thailand
Daptus vittatus Fischer von Waldheim, 1824 RU WS SS Ukraine Turkmenia Kazakhstan
Trichotichnus nishioi Habu, 1961 AM
Trichotichnus coruscus Tschitscherine, 1895 AM SH
Nipponoharpalus discrepans (A.Morawitz, 1862) AM
Parophonus (Parophonus) maculicornis (Duftsschmid, 1812) CU Georgia
Parophonus (Tachyophonus) mendax (Rossi, 1790) Azerbaijan
Parophonus (Tachyophonus) planicollis (Dejean, 1829) Azerbaijan
Parophonus (Ophonomimus) hirsutulus (Dejean, 1829) Azerbaijan Tadzhikistan
Parophonus (Ophonomimus) interstitialis (Reitter, 1900) Turkmenistan Tadzhikistan
Harpalobrachys leiroides (Motschulsky, 1844) SS AM
Harpalus cephalotes Fairmaire et Laboulbene, 1854 RU
Harpalus capito A.Morawitz, 1862 KU
Harpalus eous Tschitscherine, 1901 AM
Harpalus griseus (Panzer, 1797) RU CU WS AM Ukraine Armenia Tadzhikistan
Harpalus juriceki (Jedlicka, 1928) AM
Harpalus ussuriensis Chaudoir, 1863 AM
Harpalus rufipes (De Geer, 1774) RU CU UR WS SS MS Ukraine Georgia Azerbaijan Tadzhikistan Kazakhstan
Harpalus ronius H.Bates, 1873 AM
Harpalus tridens (A.Morawitz, 1862) AM
Harpalus tschiliensis Schauberger, 1929 AM
Harpalus coreanus Tschitscherine, 1895 AM
Harpalus paratus Casey, 1924 USA
Harpalus calceatus (Duftschmid, 1812) RU CU WS SS Ukraine Armenia Kazakhstan
Harpalus signaticornis (Duftschmid, 1812) CU SS
Harpalus tenebrosus Dejean, 1829 Ukraine Turkmenia Tadzhikistan
Harpalus litigiosus Dejean, 1829 Ukraine
Harpalus melancholicus Dejean, 1829 Moldova
Harpalus rufipalpis Sturm, 1818 France Georgia
Harpalus honestus (Duftschmid, 1812) CU Georgia
Harpalus rubripes (Duftschmid, 1812) CU WS SS MS Kazakhstan Mongolia
Harpalus attenuatus Stephens, 1828 RU
Harpalus aeneipennis (Faldermann, 1836) Georgia
Harpalus chrysopus Reitter, 1900
Harpalus chrysopus chrysopus Reitter, 1900 CU
Harpalus chrysopus abasinus Rost, 1891 Georgia
Harpalus quadripunctctus Dejean, 1829 SS CY NE Kazakhstan
Harpalus major Motschulsky, 1850 SS
Harpalus serripes (Quensel, 1806)
Harpalus serripes serripes (Quensel, 1806) RU SS Ukraine Kazakhstan
Harpalus serripes ernsti Kataev, 1995 Kazakhstan
Harpalus pseudoserripes Reitter, 1900
Harpalus pseudoserripes pseudoserripes Reitter, 1900 Kyrghyzstan Turkmenistan
Harpalus pseudoserripes apicipunctatus Mlynar Tadzhikistan
Harpalus politus Dejean, 1829 RU WS SS Kazakhstan
Harpalus flavicornis Dejean, 1829
Harpalus flavicornis flavicornis Dejean, 1829 Ukraine
Harpalus pumilus (Sturm, 1818) RU WS SS Kazakhstan
Harpalus masoreoides H.Bates, 1878 Kyrghyzstan
Harpalus picipennis Duftschmid, 1812 RU
Harpalus lutshniki Shauberger, 1932 WS SS
Harpalus pusillus Motschulsky, 1850 SS CY
Harpalus michaili Kataev, 1990 WS SS
Harpalus acupalpoides Reitter, 1900 SS
Harpalus longipalmatus Mordkovitsch, 1969 SS
Harpalus anxius (Duftschmid, 1812) RU WS SS CY Kazakhstan
Harpalus kirgisicus Motschulsky, 1844 RU WS Kazakhstan
Harpalus amariformis Motschulsky, 1844 SS
Harpalus amplicollis Ménétriès, 1848 RU SS Kazakhstan
Harpalus calathoides Motschulsky, 1844 RU SS Kazakhstan
Harpalus servus (Duftschmid, 1812) RU Kazakhstan
Harpalus subcylindricus Dejean, 1829 RU WS Kazakhstan
Harpalus egorovi Lafer, 1989 SS
Harpalus hirtipes (Panzer, 1797) RU SS Ukraine Kazakhstan
Harpalus alajensis Tschitscherine, 1898 Kyrghyzstan
Harpalus tichonis Jacobson, 1907 SS
Harpalus corporosus (Motschulsky, 1861) AM
Harpalus macronotus Tschitscherine, 1893 SS Kazakhstan
Harpalus zabroides Dejean, 1829 RU WS SS MS Ukraine Kazakhstan
Harpalus brevis Motschulsky, 1844 WS SS Kyrghyzstan Kazakhstan
Harpalus brevicornis Germar, 1824 SS CY NE
Harpalus froelichi Sturm, 1818 RU WS SS Ukraine Kyrghyzstan Kazakhstan
Harpalus flavescens (Piller et Mitterpacher, 1783) RU
Harpalus modestus Dejean, 1829 SS AM
Harpalus bungii Chaudoir, 1844 AM
Harpalus tardus (Panzer, 1797) WS
Harpalus tarsalis Mannerheim, 1825 WS SS AM
Harpalus albanicus Reitter, 1900 Kazakhstan
Harpalus latus (Linnaeus, 1758) RU WS SS AM Poland Georgia Azerbaijan Kazakhstan
Harpalus ussuricus Mlynar, 1979 AM
Harpalus torridoides Reitter, 1900 SS
Harpalus progrediens Schauberger, 1922 Ukraine
Harpalus xanthopus Gemminger et Harold, 1868 SS AM Kazakhstan
Harpalus solitaris Dejean, 1829 WS SS KU Kazakhstan
Harpalus luteicornis (Duftschmid, 1812) WS
Harpalus nigritarsis C.R.Sahlberg, 1827 SS Kazakhstan
Harpalus rufiscapus Gebler, 1833 SS
Harpalus vittatus Gebler, 1833
Harpalus vittatus vittatus Gebler, 1833 SS
Harpalus vittatus alaskensis Lindroth, 1968 NE
Harpalus udege Lafer, 1989 SS
Harpalus nigrans A.Morawitz, 1862 SS
Harpalus sinuatus Tschitscherine, 1893 SS
Harpalus fuscipalpis (Sturm, 1818) WS SS Kazakhstan
Harpalus viridanus Motschulsky, 1844 SS
Harpalus fuscicornis Ménétriès, 1832 Turkmenistan Kyrghyzstan Kazakhstan
Harpalus smaragdinus (Duftschmid, 1812) RU WS SS CY Kyrghyzstan Kazakhstan
Harpalus pallidipennis A.Morawitz, 1862 SS AM
Harpalus sarmaticus Motschulsky, 1850
Harpalus sarmaticus sarmaticus Motschulsky, 1850 RU WS SS
Harpalus sarmaticus olenini Poppius, 1906 SS
Harpalus kiritshenkoi Kataev, 1990 Kazakhstan
Harpalus cyanopterus Tschitscherine, 1897 Kazakhstan
Harpalus famelicus Tschitscherine, 1898
Harpalus famelicus fanensis Kataev et Wrase, 1993 Tadzhikistan
Harpalus pulvinatus Ménétriès, 1848 Kazakhstan
Harpalus rhemboides Solsky, 1874 Kazakhstan
Harpalus cisteloides Motschulsky, 1844
Harpalus cisteloides cisteloides Motschulsky, 1844 WS SS Kyrghyzstan
Harpalus aequicollis Motschulsky, 1844 SS
Harpalus heyrovskyi Jedlicka, 1928 SS
Harpalus optabilis Dejean, 1829 RU WS Kazakhstan
Harpalus ellipticus Ballion, 1878 SS Kazakhstan
Harpalus salinus Dejean, 1829
Harpalus salinus salinus Dejean, 1829 SS Kazakhstan
Harpalus salinus agonus Tschitscherine, 1894 Kyrghyzstan Tadzhikistan
Harpalus salinus klementzae Kataev, 1984 SS
Harpalus lumbaris Mannerheim, 1825 SS Kazakhstan
Harpalus autumnalis (Duftschmid, 1812) RU
Harpalus foveiger Tschitscherine, 1895 RU
Harpalus setiporus Reitter, 1894 Tadzhikistan
Harpalus dispar Dejean, 1829
Harpalus dispar splendens Gebler, 1830 WS SS Kazakhstan
Harpalus hospes Sturm, 1818
Harpalus hospes hospes Sturm, 1818 CU
Harpalus steveni Dejean, 1829 RU WS SS Kazakhstan
Harpalus terrestris (Motschulsky, 1844) RU Kazakhstan
Harpalus circumpunctatus Chaudoir, 1846 RU
Harpalus affinis (Schrank, 1781) RU UR WS SS MS CY AM Latvia Ukraine Georgia Kazakhstan
Harpalus tjanshanicus Semenov, 1889 Kyrghyzstan Kazakhstan
Harpalus glasunovi Kataev, 1987 Kyrghyzstan
Harpalus staudingeri Jedlicka, 1953
Harpalus staudingeri altaicus Jedlicka, 1968 Kazakhstan
Harpalus pewtzowi Tschitscherine, 1897 Kazakhstan
Harpalus bucharicus Tschitscherine, 1898 Tadzhikistan
Harpalus erosus Mannerheim, 1825 SS Kazakhstan
Harpalus obtusus (Gebler, 1833)
Harpalus obtusus obtusus (Gebler, 1833) SS
Harpalus obtusus amputatoides Mlynar, 1979 NE
Harpalus distinguendus (Duftschmid, 1812)
Harpalus distinguendus distinguendus (Duftschmid, 1812) RU WS SS MS Georgia Uzbekistan Kazakhstan
Harpalus distinguendus kidanicus Kataev, 1989 SS
Harpalus saxicola Dejean, 1829 Georgia
Harpalus oblitus Dejean, 1829
Harpalus oblitus oblitus Dejean, 1829 Kazakhstan
Harpalus smyrnensis Heyden, 1888 Armenia
Harpalus akinini Tschitscherine, 1895 RU Kazakhstan
Harpalus crates H.Bates, 1883 AM
Neophygas microcephalus (Faldermann, 1835) SS
Microderes (Microderes) brachypus (Dejean, 1829) RU Ukraine Kazakhstan
Microderes (Microderes) undulatus (Gebler, 1841) Kazakhstan
Microderes (Neopangus) namanganensis (Heyden, 1885) Kyrghyzstan
Microderes (Neopangus) intermittens (Solsky, 1874) Turkmenistan Uzbekistan
Acinopus (Acinopus) laevigatus Ménétriès, 1832 RU Ukraine Georgia Turkmenistan Kyrghyzstan Uzbekistan Tadzhikistan Kazakhstan
Acinopus (Acinopus) picipes (Olivier, 1795) France Ukraine Georgia Kyrghyzstan
Acinopus (Haplacinopus) striolatus Zoubkoff, 1833 Kazakhstan
Acinopus (Osimus) ammophilus Dejean, 1828 Turkmenistan
Acinopus (Oedematicus) megacephalus (Rossi, 1794) Azerbaijan
Ophonus (Metophonus) nitidulus Stephens, 1828 WS SS
Ophonus (Metophonus) cordatus (Duftschmid, 1812) Turkey Turkmenistan
Ophonus (Metophonus) stricticollis (Tschitscherine, 1893) AM
Ophonus (Metophonus) hystrix Reitter, 1900
Ophonus (Metophonus) hystrix hystrix Reitter, 1900 SS
Ophonus (Metophonus) hystrix dissors Tschitscherine, 1895 Kyrghyzstan
Ophonus (Metophonus) hystrix altaiensis Kataev, 2001 paratypes! SS
Ophonus (Metophonus) hystrix kamenskii Kataev et Kabak in Kataev, 2001 paratype! Kazakhstan
Ophonus (Metophonus) puncticollis (Paykull, 1798) RU WS SS Ukraine Tadzhikistan Kazakhstan
Ophonus (Metophonus) puncticeps (Stephens, 1828) CU Ukraine
Ophonus (Metophonus) rufibarbis (Fabricius, 1792) RU CU SS Moldova Ukraine Tadzhikistan Kazakhstan
Ophonus (Metophonus) melletii (Heer, 1837) Ukraine
Ophonus (Hesperophonus) similis (Dejean, 1829) Ukraine
Ophonus (Hesperophonus) azureus (Fabricius, 1775) RU CU SS Ukraine Georgia Armenia Kazakhstan
Ophonus (Hesperophonus) chlorizans Solsky, 1874 Turkmenistan Kyrghyzstan Tadzhikistan
Ophonus (Hesperophonus) subquadratus (Dejean, 1892) Ukraine Turkmenistan
Ophonus (Hesperophonus) convexicollis Ménétriès, 1832 Turkmenistan
Ophonus (Hesperophonus) cribricollis Dejean, 1829 Ukraine Turkmenistan
Ophonus (Ophonus) stictus Stephens, 1828 RU WS SS Ukraine Kazakhstan
Ophonus (Ophonus) ardosiacus (Lutshnik, 1922) Georgia Azerbaijan
Ophonus (Ophonus) diffinis (Dejean, 1829) Moldova
Ophonus (Ophonus) sabulicola (Panzer, 1796) Ukraine
Liochirus cycloderus (Solsky, 1874) Turkmenistan Kyrghyzstan
Carterus (Pristocarterus) angustus Ménétriès, 1832 Azerbaijan
Ditomus calydonius (Rossi, 1790) Turkey Turkmenistan
Dixus eremita (Dejean, 1825) RU Ukraine
Dixus obscurus (Dejean, 1825) CU Ukraine
Dixus semicylindricus Piochard de la Brulerie, 1872 Turkmenistan Uzbekistan Kyrghyzstan Kazakhstan
Eocarterus chodshenticus (Ballion, 1871)
Eocarterus chodshenticus chodshenticus (Ballion, 1871) Uzbekistan Tadzhikistan
Eocarterus esau (Heyden, 1885)
Eocarterus esau esau (Heyden, 1885) Uzbekistan
Eocarterus propagator (Reitter, 1901) Tadzhikistan
Eocarterus semenovi (Reitter, 1893) Uzbekistan
Eocarterus usgentensis (Heyden, 1884) Kazakhstan
Bronislavia robusta Semenov, 1891 Tadzhikistan
Carenochhyrus titanus Solsky, 1874 Uzbekistan
Machozethus concinnus Dohrn, 1885 Turkmenistan
Machozethus lehmanni (Ménétriès, 1849) Turkmenistan
Chilotomus chalibaeus (Faldermann, 1835) Turkmenistan
Chilotomus kuhitangi Kryzhanovskij, 1962 Turkmenistan
Chilotomus marginatus Kryzhanovskij, 1962 Turkmenistan
Amblystomus aeneolus Chaudoir Kenya


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