materials from different regions of Russia are encoded as: FS RU CU UR WS SS MS CY NE NF KM AM SH KU
Hybosorinae
Hybosorus illigeri Reiche, 1853 Turkmenistan
Tadzhikistan
Scarabaeinae
Scarabaeus sacer Linnaeus, 1758 RU
Azerbaijan Turkmenistan
Scarabaeus semipunctatus Fabricius, 1792 Spain
Scarabaeus pius Illiger, 1803 CU
Armenia Turkmenistan Tadzhikistan
Scarabaeus typhon Fischer, 1822 RU Kazakhstan
Turkmenistan
Scarabaeus carinatus Gebler, 1841 Uzbekistan
Turkmenistan Tadzhikistan
Scarabaeus babori Balthasar, 1934 Uzbekistan
Turkmenistan
Scarabaeus acuticollis Motschulsky, 1849
Turkmenistan Tadzhikistan
Scarabaeus armeniacus Menetries, 1832
CU Georgia
Scarabaeus transcaspicus Stolfa, 1938
Turkmenistan
Gymnopleurus flagellatus (Fabricius, 1787)
CU WS Azerbaijan Kazakhstan Turkmenistan
Gymnopleurus mopsus Pallas, 1781 RU SS
Ukraine Kazakhstan Turkmenistan
Gymnopleurus aciculatus Gebler, 1845 Kazakhstan
Uzbekistan Turkmenistan Tadzhikistan
Gymnopleurus sturmi MacLeay, 1821 Bulgaria
Gymnopleurus geoffroyi Fuessly, 1875 RU
UR
Gymnopleurus smaragdinus Fahraeus in Boheman, 1857 Uganda
Gymnopleurus sinuatus (Olivier, 1789) China(Taiwan)
Coprinae
Sisyphus schaefferi Linnaeus, 1758
Sisyphus schaefferi boschniaki Fischer von Waldhaim,
1823 RU UR Bulgaria
Crimea Armenia Georgia Turkmenistan
Sisyphus schaefferi morio Arrow, 1909
AM
Synapsis tmolus Fischer, 1821 Kazakhstan
Kyrghyzstan Turkmenistan Tadzhikistan
Copris lunaris (Linnaeus, 1758) RU CU
UR Latvia Crimea Kazakhstan Kyrghyzstan Turkmenistan
Copris hispanus (Linnaeus, 1764) Ukraine
Uzbekistan Turkmenistan Tadzhikistan
Copris ochus Motschulsky, 1860 AM
Copris pecuarius Lewis, 1884 AM
Copris integer Reiche, 1847 Rwanda
Copris orion Klug, 1835 Rwanda
Copris magicus Harold, Vietnam
Copris (Microcopris) reflexus F. Vietnam
Copris (Microcopris) propinquus Fels. Vietnam
Heliocopris hamadryas (Fabricius, 1775) Uganda
Onthophagus (Euonthophagus) amyntas (Olivier, 1789)
Kazakhstan Kyrghyzstan Uzbekistan Turkmenistan
Tadzhikistan
Onthophagus (Euonthophagus) gibbosus (Scriba, 1790)
Azerbaijan Kazakhstan Kyrghyzstan Uzbekistan Turkmenistan Tadzhikistan
Onthophagus (Euonthophagus) sulcicollis Reitter, 1892 Turkmenistan
Tadzhikistan Afghanistan
Onthophagus (Euonthophagus) crocatus Mulsant, 1873 Algeria
Onthophagus (Euonthophagus) dorbignyi Olsoufieff, 1900
Afghanistan
Onthophagus (Altonthophagus) sibiricus Harold, 1877
Kazakhstan Kyrghyzstan Tadzhikistan
Onthophagus (Altonthophagus) marmotae Kabakov, 1990 Tadzhikistan
Onthophagus (Altonthophagus) uniformis Heyden, 1886
AM
Onthophagus (Relictonthophagus) punctator Reitter, 1893
SS AM
Onthophagus (Gibbonthophagus) atripennis Waterhouse, 1875
AM
Onthophagus (Gibbonthophagus) fodiens Waterhouse, 1875
AM
Onthophagus (Gibbonthophagus) limbatus (Herbst)
Philippines
Onthophagus (Phanaeomorphus) japonicus Harold, 1874
AM
Onthophagus (Furconthophagus) furcatus (Fabricius, 1781) RU
CU UR Georgia Armenia Azerbaijan Turkmenistan
Onthophagus (Colobonthophagus) transcaspicus Koenig,
1888 Turkmenistan
Onthophagus (Colobonthophagus) lunatus Harold, 1868 Vietnam
Onthophagus (Onthophagus) taurus (Schreber, 1759) RU
CU Ukraine Crimea
Georgia Iran Turkmenistan Uzbekistan
Onthophagus (Onthophagus) illyricus {Scopoli, 1763) Crimea
Azerbaijan
Onthophagus (Onthophagus) bivertex Heyden, 1887
AM
Onthophagus (Palaeonthophagus) vitulus (Fabricius, 1776) RU
UR WS Kazakhstan Bulgaria
Onthophagus (Palaeonthophagus) citellorum Medvedev, 1965 RU
Onthophagus (Palaeonthophagus) quadrinodus Reitter, 1896 Uzbekistan
Onthophagus (Palaeonthophagus) gibbulus (Pallas, 1781) RU
CU UR WS SS MS AM Crimea Kazakhstan Kyrghyzstan
Onthophagus (Palaeonthophagus) laticornis Gebler, 1823
SS
Onthophagus (Palaeonthophagus) nuchicornis (Linnaeus, 1758)
RU
UR WS SS Kazakhstan
Onthophagus (Palaeonthophagus) vacca (Linnaeus, 1767) RU
CU UR Moldova Azerbaijan Turkmenistan
Onthophagus (Palaeonthophagus) lemur (Fabricius, 1781)
CU Moldova Turkmenistan
Onthophagus (Palaeonthophagus) fissicornis Krynisky, 1834
CU Armenia Azerbaijan Turkmenistan
Onthophagus (Palaeonthophagus) suturellus Brulle, 1832 RU
Azerbaijan
Onthophagus (Palaeonthophagus) conspersus Reitter, 1892 Tadzhikistan
Afghanistan
Onthophagus (Palaeonthophagus) marginalis Gebler, 1817
WS SS Kazakhstan Kyrghyzstan Turkmenistan
Uzbekistan Mongolia
Onthophagus (Palaeonthophagus) speculifer Solsky, 1876
Kazakhstan Kyrghyzstan Turkmenistan
Onthophagus (Palaeonthophagus) basipusctulatus Heyden, 1889
Turkmenistan
Onthophagus (Palaeonthophagus) pygargus Motschulsky, 1845
Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan
Tadzhikistan
Onthophagus (Palaeonthophagus) ovatus (Linnaeus, 1762) RU
CU Bulgaria Crimea
Onthophagus (Palaeonthophagus) semicornis (Panzer, 1798) RU
UR SS Kazakhstan Turkmenistan
Onthophagus (Palaeonthophagus) medvedevi Kabakov, 1982 Uzbekistan
Onthophagus (Palaeonthophagus) vlasovi Medvedev, 1958
Kazakhstan Turkmenistan
Onthophagus (Palaeonthophagus) glasunovi Koshantschikov, 1894
Tadzhikistan
Onthophagus (Palaeonthophagus) finschi Harold, 1877
Kazakhstan Kyrghyzstan Turkmenistan
Onthophagus (Palaeonthophagus) afghanus Petrovitz, 1961 Tadzhikistan
Onthophagus (Palaeonthophagus) trispinus Reitter, 1892 Turkmenistan
Tadzhikistan
Onthophagus (Palaeonthophagus) leucostigma Stevens, 1811
UR Kazakhstan Turkmenistan
Uzbekistan Tadzhikistan
Onthophagus (Palaeonthophagus) haroldi Ballion, 1871
Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan
Tadzhikistan
Onthophagus (Palaeonthophagus) truchmenus Kolenati, 1846 Azerbaijan
Turkmenistan
Onthophagus (Palaeonthophagus) ruficapillus Brulle, 1832
CU Armenia Azerbaijan
Onthophagus (Palaeonthophagus) grossepunctatus J.Muller, 1906
CU
Onthophagus (Palaeonthophagus) verticicornis (Laicharting, 1781)
Azerbaijan
Onthophagus (Palaeonthophagus) fracticornis (Preyssler, 1790)
RU
CU UR WS SS Hungary Moldova Ukraine Georgia
Armenia Kazakhstan Turkmenistan
Onthophagus (Palaeonthophagus) coenobita (Herbst, 1783)
Moldova Georgia
Onthophagus (Palaeonthophagus) angorensis Petrovitz Turkmenistan
Onthophagus (Palaeonthophagus) pseudocaccobius Reitter, 1889
Turkmenistan
Onthophagus (Palaeonthophagus) olsoufieffi Boucomont, 1924 SS
AM
Onthophagus (Palaeonthophagus) scabriusculus Harold, 1873 RU
WS SS AM
Onthophagus (Palaeonthophagus) viridis Menetries, 1832 Azerbaijan
Onthophagus (Palaeonthophagus) clitellifer Reitter, 1894 SS
Onthophagus (Palaeonthophagus) opacicollis Reitter, 1893
Bulgaria
Onthophagus (Parascatonomus) multicifrons (Endrodi, 1973) Vietnam
Onthophagus cuniculus Macley Australia
Diastellopalpus lamellicollis Qued. Rwanda
Caccobius (Caccobius) jessoensis Harold, 1867 KU
Caccobius (Caccobius) brevis Waterhouse, 1875
AM
Caccobius (Caccobius) schreberi (Linnaeus, 1767) RU
CU UR Ukraine Crimea Georgia Turkmenistan
Caccobius (Caccobius) histeroides (Menetries, 1832) RU
CU Crimea
Caccobius (Caccobius) denticollis Harold, 1897 Afghanistan
Caccobius (Caccobius) hymalayanus Jekel, 1772 Afghanistan
Caccobius (Caccophilus) christophi Harold, 1879
AM
Caccobius (Caccophilus) sordidus Harold, 1886
AM
Caccobius (Caccophilus) kelleri (Olsoufieff, 1907)
AM
Chironitis pamphilus Menetries, 1849 Uzbekistan
Turkmenistan
Chironitis eumenes (Motschulsky, 1859)
Ukraine Kazakhstan
Chironitis haroldi Ballion, 1870 Kazakhstan
Uzbekistan Turkmenistan
Chironitis moeris Pallas, 1781 RU
Kazakhstan
Chironitis sterculius Ballion, 1870 Uzbekistan
Turkmenistan
Chironitis klapperichi Ballion, 1870 Iran
Onitis damoetas Steven, 1806 Georgia
Onitis humerosus (Pallas, 1771) Kazakhstan
Uzbekistan Turkmenistan
Onitis crenatus Reiche, 1850 Rwanda
Onitis falcatus (Wulfen, 1789) Thailand
Onitis subopacus Arrow, 1931 Vietnam
Euoniticellus fulvus (Goeze, 1777)
CU UR WS Moldova Ukraine Crimea Georgia Kazakhstan
Kyrghyzstan Turkmenistan Uzbekistan Tadzhikistan
Euoniticellus pallipes (Fabricius, 1781)
RU CU Israel Turkmenistan Tadzhikistan
Aphodiinae
Aegialiini
Aegialia (Aegialia) comis (Levis, 1895) AM
Aegialia (Psammoporus) abdita (Nikritin, 1975)
SS
Aegialia (Psammoporus) friebi Balthasar, 1935 SH
Aegialia (Psammoporus) kamtschatica Motschulsky, 1860 AM
SH KM
Aegialia (Psammoporus) sabuleti Panzer, 1797
RU
Aegialia (Psammoporus) cylindrica (Eschscholtz, 1822) USA
Eremazus unistriatus Mulsant, 1851 Turkmenistan
Eremazus cribratus Semenov, 1893 Turkmenistan
Psammodini
Diastictus vulneratus Sturm, 1805 WS SS
Psammodius asper (Fabricius, 1775) SS
Psammodius (Rhyssemus) orientalis (Mulsant et Codart, 1875)
Kazakhstan Turkmenistan
Psammodius (Rhyssemus) germanus (Linnaeus, 1767) CU
WS SS Georgia Turkmenistan
Tadzhikistan
Psammodius (Rhyssemus) laevipennis Costa, 1744
Ukraine
Psammodius (Pararhyssemus) coluber Mayet, 1787 Turkmenistan
Afghanistan
Pleurophorus variolosus (Kolenati, 1846)
RU Ukraine Kazakhstan Turkmenistan Tadzhikistan
Pleurophorus caesus Creutzer, 1796 CU
Moldova Georgia Turkmenistan Tadzhikistan
Pleurophorus apicipennis Reitter, 1892
Kazakhstan Turkmenistan Tadzhikistan
Thinorycterini
Thinorycter chlamydatus Semenov et Reichardt, 1925
Turkmenistan
Aphodiini
Turanella latevittis (Reitter, 1887) Turkmenistan
Oxyomus silvestris (Scopoli, 1763) RU
CU Georgia
Heptaulacus testudinarius (Fabricius, 1775)
CU
Heptaulacus sus (Herbst, 1783) RU UR WS
Armenia
Hugary
Heptaulacus carinatus (Germar, 1824) UR
WS SS Kazakhstan
Heptaulacus villosus (Gyllenhal, 1806)
SS
Aphodius (Colobopterus) erraticus (Linnaeus, 1758) RU
UR WS SS MS Ukraine Kazakhstan Turkmenistan
Uzbekistan
Aphodius (Colobopterus) propraetor Balthasar, 1932
AM
Aphodius (Colobopterus) quadratus Reiche, 1847 KU
Aphodius (Coprimorphus) scrutator (Herbst, 1789)
Italy Hugary Crimea
Aphodius (Eupleurus) subterranius (Linnaeus, 1758) RU
CU UR WS SS CY Kazakhstan Turkmenistan
Aphodius (Eupleurus) antiquus Faldermann, 1835
SS Mongolia
Aphodius (Teuchestes) brachysomus Solsky, 1874 KU
Aphodius (Teuchestes) fossor (Linnaeus, 1758) RU
CU UR WS SS Ukraine Crimea Kazakhstan
Aphodius (Otophorus) haemorrhoidalis (Linnaeus, 1758) RU
UR WS SS AM SH KU Italy Ukraine Kazakhstan
Aphodius (Lunaphodius) lunifer Solsky, 1876
Turkmenistan Tadzhikistan
Aphodius (Ammoecius) brevis Erichson, 1848 RU
UR WS Kazakhstan
Aphodius (Aparammoecis) isaburoi Nakane, 1956 Japan
Aphodius (Bodilus) hydrohoeris (Fabricius, 1798) France
Aphodius (Bodilus) sordidus (Fabricius, 1775)
WS SS Mongolia
Aphodius (Bodilus) rufus (Moll, 1782)
UR Kazakhstan Hugary
Aphodius (Bodilus) ictericus (Laicharting, 1781) RU
CU WS SS Hungary Crimea Kazakhstan
Aphodius (Bodilus) punctipennis Erichson, 1848 WS
Kazakhstan Turkmenistan
Aphodius (Bodilus) longeciliatus Reitter, 1887
SS
Aphodius (Bodilus) lugens Creutzer, 1799 RU
CU Ukraine Crimea Kazakhstan Kyrghyzstan Turkmenistan
Tadzhikistan
Aphodius (Bodilus) sordescens Harold, 1869
WS SS Kazakhstan Mongolia
Aphodius (Bodilus) gregarius Harold, 1871 RU
Kazakhstan Mongolia
Aphodius (Bodilus) digitalis D.Koshantschikov, 1894 Turkmenistan
Aphodius (Bodilus) hastatus Reitter, 1892 Kazakhstan
Aphodius (Bodilus) strigimargo Reitter, 1892 Turkmenistan
Aphodius (Bodilus) nigriventris Reitter, 1892 Kazakhstan
Aphodius (Acanthbodilus) immundus Creutzer, 1799
CU UR WS SS Kazakhstan Turkmenistan Tadzhikistan
Aphodius (Erytus) klugi A.Schmidt, 1910
Turkmenistan
Aphodius (Erytus) aequalis A.Schmidt, 1907
Turkmenistan
Aphodius (Erytus) bucharicus Petrovitz, 1961
Turkmenistan
Aphodius (Erytus) persicus Petrovitz, 1961 Afghanistan
Aphodius (Erytus) pruinosus Reitter, 1892
Turkmenistan
Aphodius (Plagiogonus) putridus (Fourcroy, 1785) RU
UR WS SS Kazakhstan
Aphodius (Plagiogonus) praeustus Ballion, 1871 Kazakhstan
Turkmenistan
Aphodius (Pharaphodius) rugosostriatus Waterhouse, 1875 Japan
Aphodius (Acrossus) bimaculatus (Laxmann, 1770)
WS SS Kazakhstan
Aphodius (Acrossus) rufipes (Linnaeus, 1758)
WS SS AM Italy Finland Ukraine Georgia Kazakhstan
Aphodius (Acrossus) depressus (Kugelann, 1798) RU
CU UR WS SS Italy Ukraine Georgia
Aphodius (Acrossus) luridus (Fabricius, 1775) RU
CU UR WS SS Crimea Kazakhstan Kyrghyzstan
Turkmenistan Uzbekistan Tadzhikistan
Aphodius (Acrossus) planicollis Reitter, 1890
Georgia
Aphodius (Acrossus) arsenjevi Berlov, 1989
AM
Aphodius (Aganocrossus) urostigma Harold, 1862 Japan
Aphodius (Neagolius) falcispinus W.Koshantschikov, 1912
SS
Aphodius (Neagolius) haroldi D.Koshantschikov, 1894 Kazakhstan
Aphodius (Neagolius) dzhungaricus Nikolajev et Frolov, 1997
paratypes!
Kazakhstan
Aphodius (Gonaphodius) scoparius Harold, 1877 SS
Aphodius (Biralus) satellitus (Herbst, 1789) RU
Kazakhstan
Aphodius (Biralus) menetriesi Menetries, 1849
Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan
Aphodius (Agolius) makowskyi D.Koshantschikov, 1891
Kazakhstan
Aphodius (Agolius) abchasikus Reitter, 1892 Georgia
Aphodius (Agoliinus) nemoralis Erichson, 1848 RU
Belarus
Aphodius (Agoliinus) nasutus Reitter, 1887
SS
Aphodius (Agoliinus) tenebricosus A. Schmidt, 1816
WS SS
Aphodius (Aphodiellus) impunctatus Waterhouse, 1875
AM
Aphodius (Chilothorax) comma Reitter, 1892
WS SS MS Kazakhstan
Aphodius (Chilothorax) badenkoi Nikolajev, 1987
paratype! Kazakhstan
Aphodius (Chilothorax) flavimargo Reitter, 1901
Kazakhstan Turkmenistan
Aphodius (Chilothorax) distinctus (Muller, 1776) RU
CU UR WS SS MS Kazakhstan Turkmenistan Tadzhikistan
Aphodius (Chilothorax) transvolgensis Semenov, 1898 RU
WS Kazakhstan
Aphodius (Chilothorax) praenubilis Balthasar, 1938 Kyrghyzstan
Aphodius (Chilothorax) melanostictus W.Schmidt, 1840 RU
WS MS Ukraine Kazakhstan Kyrghyzstan Turkmenistan
Aphodius (Chilothorax) auliensis Balthasar, 1933 Kyrghyzstan
Aphodius (Chilothorax) clathratus Reitter, 1892 unknown
locality
Aphodius (Chilothorax) scuticollis Semenov, 1898
Kazakhstan Tadzhikistan
Aphodius (Chilothorax) sticticus (Panzer, 1798) RU
CU Belarus
Aphodius (Chilothorax) grafi Reitter, 1901
SS
Aphodius (Chilothorax) tanhensis Frolov, 2001
SS
Aphodius (Chilothorax) jakobsoni W.Koshantshikov, 1911
SS Kazakhstan
Aphodius (Chilothorax) altaicus Nikolajev, 1984 China
Aphodius (Chilothorax) exilimanus Kabakov, 1994 UR
Aphodius (Chilothorax) kerzhneri Nikolajev, 1984
WS SS Kazakhstan
Aphodius (Chilothorax) mongolaltaicus Nikolajev, 1984
SS Mongolia
Aphodius (Melinopterus) prodromus (Brahm, 1790) RU
CU UR WS SS Ukraine Latvia Kazakhstan Turkmenistan
Aphodius (Melinopterus) punctatosulcatus Sturm, 1805 RU
UR WS SS MS Kazakhstan
Aphodius (Melinopterus) sphacelatus (Panzer, 1798) RU
Aphodius (Melinopterus) circumcinctus W.Schmidt, 1846 RU
UR WS
Aphodius (Melinopterus) consputus Creutzer, 1899 RU
Aphodius (Melinopterus) scuticollis Semenov, 1898 Kazakhstan
Tadzhikistan
Aphodius (Melaphodius) caspicus Menetries, 1823 RU
WS SS MS Ukraine Kazakhstan
Aphodius (Nimbus) turkestanicus Heyden, 1881
Kazakhstan Kyrghyzstan
Aphodius (Nobius) serotinus (Panzer, 1799) RU
WS Kazakhstan
Aphodius (Nobius) circumductus Solsky, 1876 Uzbekistan
Aphodius (Nobius) korgaldzhensis Nikolajev, 1987 UR
Aphodius (Nobius) pustulifer Reitter, 1892 Uzbekistan
Aphodius (Phaeaphodius) rectus Motschulsky, 1866 UR
WS SS MS AM KU Kazakhstan China
Aphodius (Phaeaphodius) dauricus Harold, 1863 RU
WS SS MS Kazakhstan
Aphodius (Phaeaphodius) costalis Gebler, 1848
WS Kazakhstan
Aphodius (Amidorus) tomentosus (Muller, 1776)
WS SS Kazakhstan
Aphodius (Amidorus) obscurus (Fabricius, 1792) Georgia
Aphodius (Amidorus) zangi A.Schmidt, 1906 RU
WS SS Kazakhstan
Aphodius (Orodaliscus) rotundangulus Reitter, 1900 RU
WS Kazakhstan
Aphodius (Oromus) alpinus (Scopoli, 1763)
Ukraine
Aphodius (Oromus) asphaltinus Kolenati, 1846 Georgia
Aphodius (Oromus) corvinus Erichson, Belarus
Aphodius (Limarus) zenkeri (Germar, 1813) Belarus
Aphodius (Pseudacrossus) edgardi Solsky, 1876 Turkmenistan
Tadzhikistan
Aphodius (Pseudacrossus) hahni Reitter,1907 UR
Kazakhstan
Aphodius (Pseudacrossus) nasutus Reitter, 1887
SS
Aphodius (Pseudacrossus) grombczewskyi D.Koshantschikov, 1891
Kazakhstan Kyrghyzstan
Aphodius (Trichonotulus) scrofa (Fabricius, 1787)
Aphodius (Trichonotulus) scrofa scrofa (Fabricius, 1787)
Kazakhstan
Aphodius (Trichonotulus) scrofa mongolicus (Mannerhaim, 1852)
SS
Aphodius (Esymus) merdarius (Fabricius, 1775) RU
CU Turkmenistan
Aphodius (Esymus) alaiensis Reitter, 1892
Kazakhstan Turkmenistan Tadzhikistan
Aphodius (Esymus) stereotypus D.Koshantschikov, 1894 Turkmenistan
Aphodius (Mecynodes) kisilkumi Solsky, 1876
WS Turkmenistan
Aphodius (Orodalus) pusillus (Herbst, 1789) RU
CU UR WS SS MS AM KU Georgia
Kazakhstan Kyrghyzstan Turkmenistan Uzbekistan
Aphodius (Orodalus) coenosus (Panzer, 1798) RU
UR Belarus
Aphodius (Eudolus) quadriguttatus (Herbst, 1783) UR
Crimea
Turkmenistan Uzbekistan
Aphodius (Phalacronotus) quadrimaculatus (Linnaeus, 1767) Turkmenistan
Aphodius (Phalacronotus) biguttatus Germar, 1824 UR
WS Kazakhstan
Aphodius (Phalacronotus) citellorum Semenov et Medvedev, 1928
RU WS
Kazakhstan
Aphodius (Mendidius) bidens Solsky, 1876
(Kazakhstan?) Turkmenistan
Aphodius (Mendidius) sijasovi Lebedev, 1932 Kazakhstan
Aphodius (Mendidius) multiplex Reitter, 1897
WS Kazakhstan Turkmenistan
Mongolia
Aphodius (Mendidius) nelsinae (Medvedev, 1968)
Kazakhstan
Aphodius (Mendidius) repetekensis (Nikritin, 1971) Turkmenistan
Aphodius (Mendidius) ivanovi Lebedev, 1912 RU
Aphodius (Mendidaphodius) brancsiki Reitter, 1899 Turkmenistan
Aphodius (Apsteiniella) georgii (W.Koshantschikov, 1912) Turkmenistan
Aphodius (Apsteiniella) aralicus Nikolajev, 1979 paratype!
Kazakhstan
Aphodius (Loraspis) frater Mulsant et Ray, 1870 UR
WS SS Kazakhstan
Aphodius (Loraphodius) suarius Faldermann, 1835
Azerbaijan
Aphodius (Labarrus) mathhiesseni Reitter, 1907 Afghanistan
Aphodius (Aphodius) fimetarius (Linnaeus, 1758) RU
CU UR WS SS Ukraine Armenia Georgia Kazakhstan
Kyrghyzstan Turkmenistan Uzbekistan
Aphodius (Aphodius) foetens (Fabricius, 1787) RU
UR WS SS Kazakhstan
Aphodius (Aphodius) elegans Allibert, 1848 Japan
Aphodius (Agrilinus) ater (De Geer, 1774) RU
UR WS SS SH Georgia Kazakhstan
Aphodius (Agrilinus) borealis Gyllenhal, 1827 WS
AM Finland Kazakhstan
Aphodius (Agrilinus) vittatus Say, 1825
Aphodius (Agrilinus) vittatus mundus Reitter, 1893
UR SS Kazakhstan Turkmenistan
Aphodius (Agrilinus) vittatus sellatus Mannerheim, 1852
SS
Aphodius (Agrilinus) uliginosus (Hardy, 1847) RU
WS SS Belarus Kazakhstan
Aphodius (Agrilinus) lapponum Gyllenhal, 1806
SS Kazakhstan
Aphodius (Agrilinus) piceus Gyllenhal, 1808 RU
SS Kazakhstan
Aphodius (Agrilinus) uniformis Waterhouse, 1875 KU
Aphodius (Agrilinus) pratensis Nomura et Nakane KU
Aphodius (Agrilinus) isajevi Kabakov, 1994 RU
UR
Aphodius (Nialus) sturmi Harold, 1870 RU
WS Kazakhstan Turkmenistan
Aphodius (Nialus) kraatzi Harold, 1868
Kazakhstan Kyrghyzstan Turkmenistan
Aphodius (Nialus) plagiatus (Linnaeus, 1767)
WS SS Latvia Kazakhstan
Aphodius (Nialus) varians Duftschmid, 1805 RU
WS Ukraine Azerbaijan Kazakhstan Tadzhikistan
Aphodius (Calamosternus) lividus (Olivier, 1789)
AM Georgia Kazakhstan Turkmenistan
Tadzhikistan
Aphodius (Calamosternus) granarius (Linnaeus, 1767) RU
CU WS SS Georgia Azerbaijan Kazakhstan Turkmenistan
Uzbekistan Tadzhikistan
Aphodius (Calamosternus) suturifer Reitter, 1893 SS
Kazakhstan
Aphodius (Calamosternus) sublimbatus Motschulsky, 1860
AM
Aphodius (Aphodaulacus) koltzei Reitter, 1892
AM
Aphodius (Aphodaulacus) variabilis Waterhoze, 1875
AM
Melolonthinae
Melolonthini
Polyphylla (Mesopolyphylla) irrorata (Gebler, 1841)
Kazakhstan
IMAGES IN NATURE:
pict.1
Polyphylla (Mesopolyphylla) tridentata Reitter, 1890 Kyrghyzstan
Polyphylla (Xerasiobia) alba (Pallas, 1773) CU
WS Kazakhstan Turkmenistan
Polyphylla (Xerasiobia) adspersa Motschulsky, 1853 Turkmenistan
Uzbekistan Tadzhikistan Kazakhstan
Polyphylla (Polyphylla) fullo (Linnaeus, 1758) RU
Ukraine
Polyphylla (Polyphylla) olivieri (Laporte de Castelnau, 1840)
CU
Georgia Armenia
Polyphylla (Gynexophylla) laticollis Lewis, 1887 Japan
Lepidiota sp. Vietnam
Haplosternus incanus Motschulsky, 1853
AM
Melolontha (Melolontha) melolontha Linnaeus, 1758
Latvia
Melolontha (Melolontha) hippocostani Fabricius, 1801
WS SS
Melolontha (Melolontha) pectoralis Germar, 1824 CU
Georgia
Melolontha (Apropyga) clypeata Reitter, 1887
Uzbekistan
Melolontha (Apropyga) afficta Ballion, 1870
Uzbekistan
Anoxia pilosa (Fabricius, 1792) RU
Asactopholis biturberculatus Moser, Vietnam
Heptophyllini
Heptophylla picea Motschulsky, 1857 South Korea Japan
Hilyotrogus bicoloreus Heyden, 1887 AM
Toxospathius auriventris Bates, 1891 China
Rhizotroginae
Rhizotrogini
Apogonia nigroolivaceae Heyden, 1886 AM
Apogonia sp. Rwanda
Lasiopsis (Lasiopsis) caninus Zoubkov, 1829
WS SS Kazakhstan
Lasiopsis (Lasiopsis) golovjankoi Medvedev, 1951
SS AM
Lasiopsis (Lasiopsis) sahlbergi Mannerhaim, 1849
SS
Lasiopsis (Brahmina) faldermanni Kraatz, 1892
SS
Lasiopsis (Brahmina) agnellus (Faldermann, 1835)
SS
Lasiopsis (Brahmina) sedakovi (Mannerheim, 1849)
SS AM
Lasiopsis (Brahmina) duchoni Reitter, 1902
Kazakhstan
Lasiopsis (Lachnota) henningi (Fischer, 1889)
SS
Lasiopsis (Lasiexis) primaeveris (Semenov et Medvedev, 1936)
Turkmenistan
Lasiopsis (Lasiexis) dilaticollis (Ballion, 1870) Kazakhstan
Kyrghyzstan Turkmenistan
Cyphochilus insulanus Moser Taiwan
Lachnosterna sp. Rwanda
Panotrogus myschenkovi (Ballion, 1871) Kyrghyzstan
Turkmenistan Tadzhikistan
Rhizotrogus (Rhizotrogus) aestivus (Olivier, 1789) RU
Crimea Turkmenistan
Rhizotrogus (Amphimallon) altaicus (Mannerheim, 1825)
WS SS Kazakhstan
Rhizotrogus (Amphimallon) volgensis (Fischer, 1823)
Kazakhstan
Rhizotrogus (Amphimallon) solsticialis (Linnaeus, 1758)
Rhizotrogus (Amphimallon) solsticialis solsticialis (Linnaeus,
1758) Latvia
Rhizotrogus (Amphimallon) solsticialis setosus Reitter,
1902 Turkmenistan
Rhizotrogus (Amphimallon) solsticialis mesasiaticus Medvedev,
1951 Kazakhstan Kyrghyzstan
Uzbekistan Tadzhikistan
Rhizotrogus (Amphimallon) solsticialis grossatus Reitter,
1902 Crimea
Rhizotrogus (Amphimallon) solsticialis sibiricus Reitter,
1902 WS SS
Rhizotrogus (Amphimallon) irtishensis Nikolajev, 1979 paratypes!
Kazakhstan
Rhizotrogus (Amphimallon) ruficornis (Fabricius, 1775)
SS
Xanthotrogus sieversi Reitter, 1896 Turkmenistan
Xanthotrogus fortis Reitter, 1902 Uzbekistan
Holochelus fallax (Marseul, 1879) Turkmenistan
Holochelus aequinoctialis (Herbst, 1790) RU
Ukraine Kazakhstan
Holochelus tauricus (Blanchard, 1850)
Ukraine
Holochelus (Madotrogus) glabripennis (Ballion, 1871) Kyrghyzstan
Turkmenistan Uzbekistan Tadzhikistan
Holochelus (Dasitrogus) transcaspicus Brenske, 1866
Uzbekistan
Holochelus (Dasitrogus) glabricollis (Reitter, 1888) Turkmenistan
Holochelus (Dasitrogus) kirgisicus (Semenov, 1895) Kyrghyzstan
Holochelus (Dasitrogus) kuhitangicus Nikolajev, 1976 Turkmenistan
Chioneosoma (Aleucolomus) porosum (Fischer, 1824) Kazakhstan
Turkmenistan
Chioneosoma (Aleucolomus) tedschenense Semenov, 1895 Turkmenistan
Chioneosoma (Aleucolomus) vulpinum (Gyllenhal, 1817) RU
Chioneosoma (Chionotrogus) candidum (Semenov, 1902) Turkmenistan
Chioneosoma (Chionotrogus) gorilla Brenske, 1886 Turkmenistan
Chioneosoma (Chionotrogus) kizilkumenze Semenov, 1895 Kazakhstan
Turkmenistan
Chioneosoma (Chioneosoma) komarovi (Brenske, 1866) Turkmenistan
Chioneosoma (Chionotrogus) peetzi Semenov, 1898 Turkmenistan
Chioneosoma (Chionotrogus) pulvereum (Koch, 1801) RU
Chioneosoma (Chionotrogus) turcomanum Brenske, 1888 Turkmenistan
Chioneosoma (Leucolasium) baadeni Brenske, 1886 Turkmenistan
Chioneosoma (Leucolasium) rostovtzovi Semenov, 1898 Turkmenistan
Chioneosoma (Leucolasium) tschitscherini Semenov, 1895 Turkmenistan
Sophrops heydeni Brenske, 1892 AM
Sophrops planicollis Burmeister Vietnam
Sophrops cephalotes Burmeister Vietnam
Pectinichelus (Cryphaeobius) brunneus (Kraatz, 1882)
Tadzhikistan
Pectinichelus (Cryphaeobius) lopatini Medvedev, 1959
Tadzhikistan
Holotrichia sichotana (Brenske, 1896) AM
Holotrichia diomphalia (Batten, 1895) AM
China
Holotrichia inelegans (Lewis, 1895) Japan
Holotrichia lata Brenske China(Taiwan)
Holotrichia maxima Chang China
Holotrichia pruinosa Wied. Vietnam
Eotrichia titanis (Reitter, 1902) AM
Haplidia transversa (Fabricius, 1801)
Bulgaria
Trematodes tenebrioides (Pallas, 1781)
SS
Pachydeminae
Tanyproctus turanicus Reitter, 1900 Turkmenistan
Tanyproctus bucharicus Reitter, 1897 Tadzhikistan
Asiactenius tekkensis (Reitter, 1888) Turkmenistan
Asiactenius tokgajevi Medvedev, 1962 Turkmenistan
Hemictenius simplicitarsis Reitter, 1897 Tadzhikistan
Hemictenius ochripennis Reitter, 1902
Tadzhikistan
Hemictenius comatus Nikolajev, 1975 Tadzhikistan
Hemictenius lebedevi Reitter, 1908 Kazakhstan
Hemictenius lopatini Medvedev, 1959 Tadzhikistan
Hemictenius elongatus Medvedev, 1962 Tadzhikistan
Hemictenius pygmaeus Medvedev, 1962 Tadzhikistan
Hemictenius medvedevi Nikolajev, 1976 Turkmenistan
Hemictenius badkhyzicus Nikolaev, 2000 paratypes!
Turkmenistan
Hemictenius opacus (Ballion, 1870) Uzbekistan
Hemictenius latitarsis (Reitter, 1891) Kazakhstan
Turkmenistan Uzbekistan
Hemictenius magnitarsis (Reitter, 1897) Uzbekistan
Hemictenius nigrociliatus Reitter, 1897
Tadzhikistan
Hemictenius bactrianus Medvedev, 1952
Tadzhikistan
Hemictenius kryzhanovskii Nikolaev et Shukronajev, 1977 paratype!
Tadzhikistan
Sericinae
Homaloplia marginata (Fuessly, 1775) Bulgaria
Homaloplia kiritshenkoi Medvedev, 1952
Crimea
Homaloplia hirta (Gebler, 1830) RU WS
SS Kazakhstan
Homaloplia spiraea (Pallas, 1773) RU UR
WS Armenia Kazakhstan
Homaloplia nicolasi Baraud, 1965 Italy
Serica (Serica) brunnea (Linnaeus, 1758)
RU UR WS SS Ukraine Kazakhstan
Serica (Serica) ussuriensis Medvedev, 1952
AM
Serica (Serica) koltzei Reitter, 1897
AM
Serica (Ophthalmoserica) rosina Pic, 1904
AM
Serica (Ophthalmoserica) niijimai Konthwan Japan
Serica (Ophthalmoserica) karafutoensis Niijma et Kinoshita SH
KU
Serica (Trichoserica) polita (Gebler, 1832) SS
MS AM KU
Serica (Trichoserica) pilosa (Nomura) Japan
Maladera (Maladera) holosericea (Scopoli, 1772)
RU CU SS Kazakhstan
Maladera (Maladera) renardi (Ballion, 1870) SS
AM
Maladera (Aserica) punctatissima (Faldermann, 1835)
Moldova Turkmenistan
Maladera (Aserica) euphorbia (Burmeister, 1895) Kazakhstan
Turkmenistan Tadzhikistan
Maladera (Aserica) lukjanovitschi (Medvedev, 1966) SS
Maladera (Aserica) orientalis (Motschulsky, 1857)
AM
Maladera (Aserica) castanea Arrow, 1913 SS
AM Georgia
Maladera (Aserica) matrida Argamon, 1986
Israel
Maladera (Aserica) spissigrada (Brenske, 1897)
AM
Leucoserica arenicola (Solsky, 1876) Turkmenistan
Sericania fuscolineata Motschulsky, 1860
AM SH
Sericania sachalinensis Matsumura, 1911
SH KU
Triodonta aquila (Castelnau, 1840) France
Hopliinae
Ectinohoplia rufipes (Motschulsky, 1860)
AM SH KU
Ectinohoplia obducta (Motschulsky, 1857)
Japan Taiwan
Ectinohoplia variabilis Reitter, 1903 Taiwan
Hoplia (Decamera) cincticollis (Faldermann, 1833) SS
Mongolia
Hoplia (Decamera) hauseri Reitter, 1904 Kazakhstan
Kyrghyzstan
Hoplia (Decamera) djukini Jacobson, 1914
AM
Hoplia (Hoplia) parvula Krynicky, 1832
RU
Hoplia (Hoplia) zaitzevi Jacobson, 1914
RU
Hoplia (Hoplia) detrita Solsky, 1876 RU
Hoplia (Hoplia) farinosa Linnaeus, 1761
Bavaria
Hoplia (Hoplia) graminicola Fabricius, 1792
Georgia
Hoplia (Hoplia) brevipes Medvedev, 1951
CU
Hoplia (Hoplia) pallinosa Krynicky, 1832
AM Georgia
Hoplia (Hoplia) praticola Duftschmidt, 1805
Bulgaria
Hoplia (Euchromoplia) aureola (Pallas, 1781) SS
Hoplia (Euchromorplia) communis Waterhouse, 1875 Japan
Hoplia ?elegantula Whit. Japan
Hoplia sp. China(Taiwan)
Cetoniinae
Trichiini
Gnorimus subopacus Motschulsky, 1860 SS AM
Gnorimus viridiopacus (Lewis, 1887) KU
Gnorimus octopunctatus Fabricius, 1775
RU
Gnorimus miwai (Chujo, 1941) China(Taiwan)
Lasiotrichius succinctus (Pallas, 1781)
SS AM China
Trichius orientalis Reitter, 1894 Crimea
Trichius fasciatus (Linnaeus, 1787) RU
CU UR WS SS AM SH KM Finland Latvia Ukraine Kazakhstan
Finland
Osmoderma barnabita Motschulsky, 1860
AM
Osmoderma eremita Scopoly, 1763 RU
Goliathini
Rhomborrhina japonica (Hope, 1841) Japan
Rhomborrhina unicolor Motschulsky, 1861 Japan
Rhomborrhina polita Waterhoose, 1875 Japan
Trigonophorus rothschildi Fairmaire, 1891
Trigonophorus rothschildi varians (Bourgoin, 1914) China(Taiwan)
Stephanorrhina sp. Rwanda
Eudicella smithi (MacLeay, 1838) Rwanda
Diplognatha gagates (Forster, 1771) Rwanda
Gymnetini
Cotinis nitida (Linnaeus, 1764) USA
Coelodera penicillata (Hope, 1831)
Coelodera penicillata formosana China(Taiwan)
Euselatos proxima China(Taiwan)
Clynteria ducalis davidis Fairmaere China(Taiwan)
Clynteria 14-maculata (F.) Sri Lanka
Cetoniini
Epicometis (Epicometis) hirta (Poda, 1761)
RU CU Crimea
Epicometis (Epicometis) hirtiformis Reitter, 1913 Kazakhstan
Kyrghyzstan
Epicometis (Epicometis) spinifrons (Reitter, 1889) Turkmenistan
Epicometis (Epicometis) turanica (Reitter, 1888) Turkmenistan
Epicometis (Epicometis) suturalis Reitter, 1913
Armenia
Epicometis (Epicometis) senicula (Menetries, 1832)
Armenia
Heterocnemis graeca (Brulle, 1832) Greece
Oxythyrea funesta (Poda, 1761) RU CU UR
WS SS Crimea Kazakhstan
Bulgaria
Oxythyrea albopicta (Motschulsky, 1845)
Uzbekistan
Oxythyrea cinctella (Schaum, 1841) Armenia
Kyrghyzstan Kazakhstan Turkmenistan UzbekistanTadzhikistan
Stalagmopygus albella (Pallas, 1781) Turkmenistan
Tadzhikistan Afghanistan
Aethiessa szekessyi Brasavola, 1939 Tadzhikistan
Aethiessa mesopotamica Burmeister, 1842
Aethiessa mesopotamica rugipennis Burmeister, 1842 Turkmenistan
Cetonia (Eucetonia) magnifica Ballion, 1870
SS AM China
Cetonia (Eucetonia) viridiopaca (Motschulsky, 1860)
SS CY AM
Cetonia (Eucetonia) roelofsi Harold, 1880
KU Japan
Cetonia (Cetonia) aurata (Linnaeus, 1761)
Cetonia (Cetonia) aurata aurata (Linnaeus, 1761)
RU Latvia Belarus Ukraine Crimea Bulgaria
Slovenia
Cetonia (Cetonia) aurata viridiventris Reitter, 1896
UR WS SS KU Kazakhstan Kyrghyzstan
Cetonia (Cetonia) aurata pisana Heer, 1891
Italy
Cetonia (Cetonia) aurata pallida (Drury, 1770)
CU Armenia Georgia
Cetonia (Cetonia) aeratula Reitter, 1891
Azerbaijan
Protaetia (Liocola) lugubris (Herbst, 1786)
Protaetia (Liocola) lugubris lugubris (Herbst, 1786)
RU WS SS Hungary Kazakhstan
Protaetia (Liocola) lugubris orientalis Medvedev, 1964
AM
Protaetia (Liocola) insperata (Lewis, 1879)
KU
Protaetia (Calopotosia) mandschuriensis Smurhoff, 1933
AM
Protaetia (Calopotosia) brevitarsis (Lewis, 1879)
AM
Protaetia (Cetonischema) aeruginosa (Drury, 1770)
RU
Protaetia (Eupotosia) affinis (Andersch, 1797)
RU
Protaetia (Potosia) cuprea (Fabricius, 1775)
Protaetia (Potosia) cuprea cuprea (Fabricius, 1775)
France Italy
Protaetia (Potosia) cuprea obscura (Andersch, 1797)
Bulgaria
Protaetia (Potosia) metallica (Herbst, 1782)
Protaetia (Potosia) metallica metallica (Herbst, 1782)
RU UR WS SS Finland Latvia Ukraine Kazakhstan
Protaetia (Potosia) metallica daurica (Motschulsky, 1860)
SS AM
Protaetia (Potosia) cuprina (Motschulsky, 1849)
CU Crimea
Protaetia (Potosia) hieroglyphica (Menetries, 1832)
Azerbaijan Turkmenistan
Protaetia (Potosia) marginicollis (Ballion, 1870) Kazakhstan
Turkmenistan Uzbekistan Tadzhikistan
Protaetia (Potosia) angustata (Germar, 1817) Croatia
Protaetia (Potosia) lenzi (Harold) Japan
Protaetia (Netocia) funebris (Gory et Percheron, 1833)
Turkmenistan
Protaetia (Netocia) turkestanica (Kraatz, 1886)
Protaetia (Netocia) turkestanica turkestanica (Kraatz,
1886) Turkmenistan
Protaetia (Netocia) turkestanica cyanea (Kraatz, 1886)
Turkmenistan Uzbekistan Tadzhikistan
Protaetia (Netocia) bogdanovi (Solsky, 1875)
Kyrghyzstan Tadzhikistan
Protaetia (Netocia) trojana (Gory et Percheron, 1833)
Armenia Georgia
Protaetia (Netocia) prototricha (Fischer, 1842) Kazakhstan
Protaetia (Netocia) hungarica (Herbst, 1790)
Protaetia (Netocia) hungarica hungarica (Herbst, 1790)
CU WS Latvia
Protaetia (Netocia) hungarica armeniaca (Menetries, 1832)
Armenia
Protaetia (Netocia) hungarica pseudoviridana Medvedev, 1947
Turkmenistan
Protaetia (Netocia) hungarica inderiensis (Krynicky, 1832) Kazakhstan
Protaetia (Netocia) hungarica sibirica (Gebler, 1829)
SS Mongolia
Protaetia (Netocia) persica (Kraatz, 1886)
Turkmenistan
Protaetia (Netocia) excavata (Gory et Percheron, 1833)
Turkmenistan
Protaetia (Netocia) interruptocostata (Ballion, 1870) Kyrghyzstan
Tadzhikistan
Protaetia (Netocia) karelini (Zoubkov, 1829)
Protaetia (Netocia) karelini karelini (Zoubkov, 1829) UR
Kazakhstan
Protaetia (Netocia) karelini agglomerata (Solsky, 1876)
Turkmenistan Uzbekistan
Protaetia (Netocia) morio (Fabricius, 1781)
Italy
Protaetia (Netocia) vidua (Gory & Percheron, 1833) Bulgaria
Protaetia (Netocia) asiatica (Faldermann, 1835)
Azerbaijan
Protaetia (Pseudonetocia) kulabensis Reitter, 1893 Tadzhikistan
Protaetia (Protaetia) orientalis (Gory & Percheron, 1833)
Japan
Protaetia culta Waterhouse 1879 Japan
Bricoptis variolosa (Gory & Percheron, 1833) Madagascar
Poecilophilides rusticola (Burmeister, 1842)
AM Japan
Glycyphana fulvistemma (Motschulsky, 1860)
AM China(Taiwan)
Glycyphana nepalensis
Glycyphana nepalensis roguesi Bourgoin Vietnam
Glycyphana georgievici Miksic Philippines
Glycyphana incongrua Janson, 1878 China(Taiwan)
Glycyphana pygmaea Mohnike Malaysia
Glycyphana cuculus Burmeister, 1842 Philippines
Oxycetonia jucunda (Faldermann, 1835)
AM China
Oxycetonia forticula (Janson) Japan
Oxycetonia histrio (Olivier) Thailand
Pachnoda sinuata (Fabricius, 1775) Rwanda
Pachnoda peregrina marginata (Drury, 1773) Uganda
laboratory
???ini
Anochilia nitida Pouillaude, 1915 Madagascar
Cremastochilini
Campsiura trguttulata Mohnike Indonesia
Clinterocera scabrosa Motschulsky, 1853 China
Valgini
Valgus hemipterus (Linnaeus, 1758) RU CU
Crimea Kyrghyzstan
Dasyvalgus angusticollis (Waterhose, 1875) Japan
Dasyvalgus montivagus Moser Vietnam
Idiovalgus curtus Paulian, 1961 Vietnam
Idiovalgus ater Pic Vietnam
Dynastinae
Dynastini
Allomyrina dichotomus (Linnaeus, 1758) Japan
Phileurini
Eophileurus chinensis (Faldermann, 1835) Japan
Oryctini
Oryctes (Oryctes) nasicornis (Linnaeus, 1758)
Oryctes (Oryctes) nasicornis nasicornis (Linnaeus, 1758)
RU CU WS SS Ukraine Crimea Armenia Kazakhstan
Italy
Oryctes (Oryctes) nasicornis punctipennis Motschulsky, 1860
Tadzhikistan
Oryctes (Eremoryctes) ata Semenov et Medvedev, 1932
Turkmenistan
Oryctes (Howoryctes) sjostedti Kolbe Rwanda
Orizabrus clunalis (LeConte, 1856) USA
Strategus aloeus (Linnaeus, 1758) Cuba
Pentodon idiota (Herbst, 1798) CU Ukraine
Crimea Iraq
Pentodon minutum Reitter, 1887 Turkmenistan
Pentodon bidens (Pallas, 1771)
Pentodon bidens bidens (Pallas, 1771)
Azerbaijan Kazakhstan
Turkmenistan Uzbekistan Tadzhikistan
Pentodon bidens punctatum Villers, 1899
Italy
Pentodon algerinum
Pentodon algerinum bispinifrons Reitter, 1894 Tadzhikistan
Pentodon quadridens (Gebler, 1845)
Pentodon quadridens quadridens (Gebler, 1845) Kazakhstan
Pentodon quadridens patruelis Frivaldszky, 1890 Mongolia
Pentodon sulcifrons Kuster, 1848 CU
Azerbaijan
Podalgus cuniculus Burmeister, 1847
Podalgus cuniculus infantulus (Semenov, 1890)
Turkmenistan
Phyllognathus excavatus (Forster, 1771) (=hauseri Reitter,
1894) Israel Turkmenistan Tadzhikistan
Eutyctus deserti Semenov, 1890 Turkmenistan
Pycnoschema diversum Peringuey, 1901 Rwanda
Xylotrupes gideon (Linnaeus)
Xylotrupes gideon australicus (Thomson) Australia
Rutelinae
Rutelini
Callistethus waterstraati ???
Callistethus
waterstraati bawangensis Wada, 2000 paratype!
Indonesia
Parastasia nigriceps Westwood, 1841 Indonesia
Peltonotus
sulawesiensis Wada, 1990 paratype!
Indonesia
Anomalini
Pharaonus lederi Reitter, 1888 Turkmenistan
Pharaonus semenovi Reitter, 1887 Turkmenistan
Phyllopertha diversa Waterhouse, 1875 Japan
Phyllopertha horticola (Linnaeus, 1758)
RU CU WS SS AM Italy Kazakhstan China
Phyllopertha irregularis Waterhouse, 1875 Japan
Phyllopertha intermixta (Arrow, 1913)
KU Japan
Phyllopertha arinicollis Ohaus, 1905 Vietnam
Phyllopertha ?humeralis Fairmaire China
Pseudosinghala dalmanni (Gyllenhal, ) Vietnam
Pseudosinghala transversa Burmaister Vietnam
Rhombonyx holosericea (Fabricius, 1787)
RU WS SS SH
Rhombonyx pekinensis Heyden, 1886 China
Rhombonyx testaceipes (Motschulsky, 1860)
Japan
Rhombonyx ussuriensis Medvedev, 1951 AM
Popillia atrocoerulea Bates, 1888 AM
Popillia indigonacea Motschulsky, 1853
AM China South Korea
Popillia quadriguttata (Fabricius, 1787)
Popillia quadriguttata quadriguttata (Fabricius, 1787)
AM
Popillia quadriguttata japonica (Newmann, 1838)
KU USA Japan
Popillia cyanea (Hope, 1831) Vietnam
Proagopertha lucidula (Faldermann, 1835)
AM
Mimela flavilabris Waterhose, 1875 KU
Japan
Mimela inonea Nomura Japan
Anomala (Euchronomala) cuprea (Hope, 1839)
KU
Anomala (Euchronomala) viridana Kolbe, 1886
AM
Anomala (Euchronomala) mongolica Faldermann, 1835
AM
Anomala (Euchronomala) albopilosa Hope, 1839 Japan
Anomala edentula Ohaus Japan
Anomala (Anomala) vitis (Fabricius, 1775) Hungary
Anomala (Anomala) dubia (Scopoli, 1763)
RU CU Latvia
Anomala (Anomala) abchasica Motschulsky, 1853
CU Georgia
Anomala (Anomala) lucens Ballion, 1871
SS KU
Anomala (Anomala) daimiana Harold, 1877
Japan
Anomala (Anomala) rufocuprea Motschulsky, 1860
KU Japan
Anomala (Anomala) motschulskyi Harold, 1877
Japan
Anomala (Anomala) ogloblini Medvedev, 1951
AM
Anomala (Anomala) luculenta Erichson, 1847
SS AM
Anomala (Anomala) cupripes Hope Vietnam
Anomala (Anomala) antiqua Gyllenhal, 1817 Vietnam
Anomala (Psammoscaphaeus) vittata Gebler, 1841 Kazakhstan
Anomala (Psammoscaphaeus) metonidia Reitter, 1903 Kyrghyzstan
Anomala (Psammoscaphaeus) errans (Fabricius, 1775)
RU Crimea Ukraine
Anomala (Psammoscaphaeus) oxiana Semenov, 1891 Tadzhikistan
Anomala dimidiata (Hope, 1831) Afghanistan
Anomala barbata Burmeister Afghanistan
Anomala costata Hope Japan
Trigonocnemis (Ammogenia) varentzovi Semenov, 1895
Turkmenistan
Blitopertha (Exomala) pallidipennis Reitter, 1903
AM
Blitopertha (Exomala) orientalis Waterhose, 1875
Japan
Blitopertha (Blitopertha) variabilis Ballion, 1870 Kazakhstan
Turkmenistan Uzbekistan Tadzhikistan
Blitopertha (Blitopertha) lineata Fabricius, 1798
RU Armenia Azerbaijan Turkmenistan
Blitopertha (Blitopertha) conspurcata Harold, 1878
AM
Blitopertha (Asiopertha) arenicola Mulsant, 1870
CU
Cyriopertha (Cyrioperta) glabra (Gebler, 1841) Kazakhstan
Uzbekistan Tadzhikistan
Cyriopertha (Cyrioperta) arcuata (Gebler, 1832)
SS
Cyriopertha (Megapertha) massageta Kirsch, 1881
Turkmenistan Tadzhikistan
Anisoplia (Chaetopteroplia) leucaspis Laporte, 1840
Turkmenistan
Anisoplia (Chaetopteroplia) segetum (Herbst, 1783)
Anisoplia (Chaetopteroplia) segetum segetum (Herbst, 1783)
RU Bulgaria
Anisoplia (Chaetopteroplia) segetum zoubkovi Krynicky,
1832 Kazakhstan
Anisoplia (Chaetopteroplia) syriaca Burmeister, 1844 Crimea
Anisoplia (Autanisoplia) austriaca (Herbst, 1783)
RU Ukraine
Anisoplia (Lasioplia) campicola Menetries, 1832
RU UR WS SS Kazakhstan
Anisoplia (Lasioplia) koenigi Reitter, 1895
Turkmenistan
Anisoplia (Lasioplia) signata Faldermann, 1835 Armenia
Georgia
Anisoplia (Lasioplia) brenskei Reitter, 1889
RU Georgia
Anisoplia (Anisoplia) farraria Erichson, 1847 Armenia
Anisoplia (Anisoplia) turcomana Zaitzev, 1917
Turkmenistan
Anisoplia (Anisoplia) agricola (Poda, 1761)
RU CU WS Kazakhstan
Anisoplia (Anisoplia) zwicki Fischer, 1823
RU
Anisoplia (Ammanisoplia) deserticola Fischer, 1823
RU WS
Parastasia nigriceps Westwood, 1841 Philippines
Adoretini
Phaeadoretus comptus (Menetries, 1849) Kazakhstan
Turkmenistan Uzbekistan
Adoretus pruinosus Ballion, 1870 Kazakhstan
Adoretus nigrifrons (Steven, 1809) RU
CU Turkmenistan Uzbekistan Kyrghyzstan
Tadzhikistan
Adoretus discolor (Faldermann, 1835) Armenia
Adoretus rufifrons Reitter, 1903 Mongolia
Adoretus convexus Burmeister, Vietnam
Adoretus sinicus Burmeister, 1844 Vietnam
Adoretus (Lepadoretus) tenuimaculatus Waterhouse, 1875
Pseudoadoretus validus Semenov, 1889 Turkmenistan
Pseudoadoretus dilutellus Semenov, 1889
Turkmenistan
Pseudoadoretus phthisicus Dohrn, 1882
Uzbekistan